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In [3]:
from bs4 import BeautifulSoup
In [72]:
import os, shutil, re, string
In [5]:
soup = BeautifulSoup (open("1124-G-1-layout.html"))
In [6]:
print(soup.prettify())
<!DOCTYPE html PUBLIC "-//W3C//DTD XHTML 1.0 Transitional//EN" "http://www.w3.org/TR/xhtml1/DTD/xhtml1-transitional.dtd">
<html lang="bg" xml:lang="bg" xmlns="http://www.w3.org/1999/xhtml" xmlns:php="http://php.net/xsl" xmlns:tp="http://www.plazi.org/taxpub" xmlns:xlink="http://www.w3.org/1999/xlink">
<head xmlns="">
<title>
A new endogean, anophthalmous species of Parazuphium Jeannel from Northern Morocco (Coleoptera, Carabidae), with new molecular data for the tribe Zuphiini
</title>
<meta content="text/html; charset=utf-8" http-equiv="Content-Type"/>
<meta content="" name="description"/>
<meta content="" name="keywords"/>
<meta content="global" name="distribution"/>
<meta content="index, follow, all" name="robots"/>
<script language="javascript">
if(self.location==top.location){
document.write('<link rel="stylesheet" href="http://www.pensoft.net/J_FILES/img/xsl-large.css" media="all" title="default">');
}else{
document.write('<link rel="stylesheet" href="http://www.pensoft.net/J_FILES/img/xsl.css" media="all" title="default">');
}
</script>
<script type="text/javascript">
function swapDivClassname(pLink, pId, pClassName){
var lDiv = document.getElementById(pId);
if( lDiv ){
if( lDiv.className == pClassName ){
lDiv.className = '';
pLink.className = 'inactive'
}else{
lDiv.className = pClassName;
pLink.className = ''
}
}
}
function openw(url, title, options) {
var newwin = window.open(url, title, options);
newwin.focus();
}
function showBaloon(pBaloonType, pId, pEvent) {
if( !pId )
return;
if( !gBaloon || !gBaloonWrapper)
return;
pEvent = pEvent || window.event;
clearTimeout(gTimeout);
if( gActiveBaloonId && gActiveBaloonType == pBaloonType && gActiveBaloonId != pId ){
gFlagToHideBaloon = true;
hideBaloon(gActiveBaloonId);
}else if(gActiveBaloonType == pBaloonType && gActiveBaloonId == pId ){
return;
}
gActiveBaloonId = pId;
gActiveBaloonType = pBaloonType;
gFlagToHideBaloon = false;
var lEventTarget = pEvent.target || pEvent.srcElement;
if( pBaloonType == 1 ){//xref
var lReference = document.getElementById('reference_' + pId);
var lText = 'Reference not found!';
if( lReference ){
lText = lReference.innerHTML;// || lReference.innerText;
}
gBaloon.innerHTML = lText;
gBaloonInnerWrapper.className = 'largeBaloon';
}else if( pBaloonType == 2 ){//Taxoni
var lReferenceName = 'taxon_' + pId + '_baloon_div';
var lReference = document.getElementById(lReferenceName);
gBaloonInnerWrapper.className = 'smallBaloon';
if( lReference ){
lText = lReference.innerHTML;// || lReference.innerText;
}else{
lText = 'Error during loading taxa data';
}
gBaloon.innerHTML = lText;
}else if( pBaloonType == 3 ){//Figuri
var lFigure = document.getElementById('fig_' + pId);
var lText = 'Figure not found!';
if( lFigure ){
lText = lFigure.innerHTML;// || lFigure.innerText;
}
gBaloon.innerHTML = lText;
gBaloonInnerWrapper.className = 'largeBaloon';
}else if( pBaloonType == 4 ){//Tablici
var lTable = document.getElementById('table_' + pId);
var lText = 'Table not found!';
if( lTable ){
lText = lTable.innerHTML;// || lTable.innerText;
}
gBaloon.innerHTML = lText;
gBaloonInnerWrapper.className = 'largeBaloon';
}
gActiveBaloonId = pId;
displayBaloon(lEventTarget, pEvent);
}
function displayBaloon(pEventTarget, pEvent){//Pozicionira i pokazva balona
if( !gBaloon || !gBaloonWrapper)
return;
var lUseWindowMaxY = parseInt('0');
gBaloonWrapper.style.display="block";
positionBaloon(pEventTarget, pEvent, lUseWindowMaxY);
}
function hideBaloonEvent(pId, pEvent){
if( !gBaloon || !gBaloonWrapper)
return;
gFlagToHideBaloon = true;
gTimeout = setTimeout("hideBaloon('" + pId + "')", 10);
}
function hideBaloon(pId){
clearTimeout(gTimeout);
if( !gFlagToHideBaloon )
return;
gBaloonIsShown = false;
gActiveBaloonId = 0;
gActiveBaloonType = false;
gBaloonWrapper.style.display="none";
}
function initBalloons(){
gBaloon = document.getElementById('baloonContent');
gBaloonIsShown = false;
gActiveBaloonId = 0;
gTimeout = false;
gActiveBaloonType = false;
gBaloonWrapper = document.getElementById('baloonWrapper');
gBaloonInnerWrapper = document.getElementById('baloonInnerWrapper');
gBaloonArrowImg = document.getElementById('baloonArrowImg');
}
function positionBaloon(pEventTarget, pEvent, pUseWindowMaxY){
var lMousePos = getMousePosition(pEvent);
var lContentWrapper = document.getElementById('contentWrapper');
//var lY = pEventTarget.offsetTop + pEventTarget.offsetHeight;
var lY = pEventTarget.offsetHeight;
//var lX = pEventTarget.offsetLeft;
var lX = 0;
var lParent = pEventTarget;
do {
lY += lParent.offsetTop;
lX += lParent.offsetLeft;
//alert(lParent.tagName + '-' + lY);
}
while((lParent = lParent.offsetParent) && lParent.id != 'contentWrapper')
/*var lXMax = lMousePos['xMax'];*/
var lXMax = lContentWrapper.offsetWidth - 20;//-20 zaradi scrolla
var lYMax;
if( pUseWindowMaxY ){
lYMax = lMousePos['yMax'];
}else{
lYMax = lContentWrapper.scrollHeight;
}
lYMax = lYMax - 40;
var lClassName = 'baloonWrapper';
var lLeft = 0;
var lTop = 0;
var lImageName = 'http://www.pensoft.net/J_FILES/img/ext_details/baloon_arrow';
if( lY + gBaloonWrapper.offsetHeight <= lYMax ){
lTop = lY;
lClassName = lClassName + 'Bottom';
lImageName = lImageName + '_bottom';
}else{
lTop = lY - gBaloonWrapper.offsetHeight - pEventTarget.offsetHeight;
if( lTop < 0 )
lTop = 0;
lClassName = lClassName + 'Top';
lImageName = lImageName + '_top';
}
if( lX + gBaloonWrapper.offsetWidth <= lXMax ){
lLeft = lX;
lClassName = lClassName + 'Right';
lImageName = lImageName + '_right';
}else{
lLeft = lX -gBaloonWrapper.offsetWidth + 50;
if( lLeft < 0 )
lLeft = 0;
lClassName = lClassName + 'Left';
lImageName = lImageName + '_left';
}
lImageName = lImageName + '.gif';
gBaloonWrapper.style.left = lLeft + 'px';
gBaloonWrapper.style.top = lTop + 'px';
gBaloonWrapper.className = lClassName;
gBaloonArrowImg.setAttribute('src', lImageName);
}
function submitForm( pFormName ){
eval('document.' + pFormName + '.submit()');
}
function showLocalitiesMenu(){
var lMenu = document.getElementById('localitiesMenuDiv');
if( lMenu )
lMenu.style.display='block';
}
function hideLocalitiesMenu(){
var lMenu = document.getElementById('localitiesMenuDiv');
if( lMenu )
lMenu.style.display='none';
}
function getMousePosition(pEvent){
if (document.layers) {
// When the page scrolls in Netscape, the event's mouse position
// reflects the absolute position on the screen. innerHight/Width
// is the position from the top/left of the screen that the user is
// looking at. pageX/YOffset is the amount that the user has
// scrolled into the page. So the values will be in relation to
// each other as the total offsets into the page, no matter if
// the user has scrolled or not.
xMousePos = pEvent.clientX;
yMousePos = pEvent.clientY;
xMousePosMax = window.innerWidth + window.pageXOffset;
yMousePosMax = window.innerHeight + window.pageYOffset;
} else if (document.all) {
// When the page scrolls in IE, the event's mouse position
// reflects the position from the top/left of the screen the
// user is looking at. scrollLeft/Top is the amount the user
// has scrolled into the page. clientWidth/Height is the height/
// width of the current page the user is looking at. So, to be
// consistent with Netscape (above), add the scroll offsets to
// both so we end up with an absolute value on the page, no
// matter if the user has scrolled or not. ;
xMousePos = window.event.x + document.body.scrollLeft;
yMousePos = window.event.y + document.body.scrollTop;
xMousePosMax = document.body.clientWidth + document.body.scrollLeft;
yMousePosMax = document.body.clientHeight + document.body.scrollTop;
} else if (document.getElementById) {
// Netscape 6 behaves the same as Netscape 4 in this regard
xMousePos = pEvent.clientX;
yMousePos = pEvent.clientY;
xMousePosMax = window.innerWidth + window.pageXOffset;
yMousePosMax = window.innerHeight + window.pageYOffset;
}
return {
'x' : xMousePos,
'y' : yMousePos,
'xMax' : xMousePosMax,
'yMax' : yMousePosMax
};
}
var gHolderDiv;
var gScrolDiv;
var gRightBtn;
var gLeftBtn;
var gLinksTable;
var gMinLeft;
function initScrollConstants(){
gHolderDiv = document.getElementById('scrollHolderDiv');
gScrolDiv = document.getElementById('scrollDiv');
gRightBtn = document.getElementById('scrollRightBtn');
gLeftBtn = document.getElementById('scrollLeftBtn');
gLinksTable = document.getElementById('linksButtonHolder');
gScrolDiv.style.width = gLinksTable.offsetWidth + 'px';
gHolderDiv.style.width = '';
var gHolderWidth = gHolderDiv.offsetWidth;
var gScrolWidth = gScrolDiv.offsetWidth;
gMinLeft = gHolderWidth - gScrolWidth;
if(gMinLeft > 0)
gMinLeft = 0;
gCurrentLeftPos = 0;
gInterval = null;
changeDivPosition(gCurrentLeftPos);
}
function scrollDivRight(){
if( gMinLeft >= 0 ){
stopScroll()
return;
}
if( gCurrentLeftPos - 1 > gMinLeft )
gCurrentLeftPos = gCurrentLeftPos - 1;
else{
stopScroll()
gCurrentLeftPos = gMinLeft;
}
changeDivPosition(gCurrentLeftPos);
}
function scrollDivLeft(){
if( gMinLeft >= 0 ){
stopScroll()
return;
}
if( gCurrentLeftPos + 1 < 0 )
gCurrentLeftPos = gCurrentLeftPos + 1;
else{
gCurrentLeftPos = 0;
stopScroll()
}
changeDivPosition(gCurrentLeftPos);
}
function checkBtnClasses(){
if( gLeftBtn ){
if( gCurrentLeftPos == 0 || gMinLeft >= 0 ){
gLeftBtn.className = 'LeftScrollBtnLinkInactive';
}else{
gLeftBtn.className = 'LeftScrollBtnLinkActive';
}
}
if( gRightBtn ){
if( gCurrentLeftPos == gMinLeft || gMinLeft >= 0 ){
gRightBtn.className = 'RightScrollBtnLinkInactive';
}else{
gRightBtn.className = 'RightScrollBtnLinkActive';
}
}
}
function changeDivPosition(pLeft){
gScrolDiv.style.left = pLeft + 'px';
checkBtnClasses()
}
function stopScroll(){
if(gInterval)
clearInterval(gInterval)
}
function startScroll(pDir){
if( gMinLeft >= 0 ){
return;
}
if( pDir == 0){//Right
gInterval = setInterval("scrollDivRight('right')", 20);
}else{
gInterval = setInterval("scrollDivLeft('left')", 20);
}
}
function getfnlink(pFn, pEl) {
lDiv = document.getElementById(pFn);
lEl = lDiv.firstChild;
while (lEl.nodeType != 1) {
lEl = lEl.nextSibling;
}
for (var i = 0; i < lEl.childNodes.length; i++) {
if (lEl.childNodes[i].tagName && lEl.childNodes[i].tagName.toLowerCase() == 'a') {
lHref = lEl.childNodes[i].href;
break;
}
}
if (lHref)
pEl.href = lHref;
}
</script>
</head>
<body>
<div class="" id="highlightWrapper" xmlns="">
<div class="highlightTaxonNameWrapper" id="highlightTaxonNameWrapper">
<div class="highlightTaxonTreatmentWrapper" id="highlightTaxonTreatmentWrapper">
<div class="highlightCountryWrapper" id="highlightCountryWrapper">
<div class="highlightCitationWrapper" id="highlightCitationWrapper">
<div class="highlightTypeescWrapper" id="highlightTypeescWrapper">
<div class="highlightCollectionWrapper" id="highlightCollectionWrapper">
<div class="highlightLocalityWrapper" id="highlightLocalityWrapper">
<div class="wrapper">
<div class="headWrapper">
<table cellpadding="0" cellspacing="1" class="highlightButtonHolder" width="100%">
<tbody>
<tr>
<td class="taxon-treatmentsButton">
<a href="#" onclick="swapDivClassname(this, 'highlightTaxonTreatmentWrapper', 'highlightTaxonTreatmentWrapper')">
Taxon treatments
</a>
</td>
<td class="taxon-namesButton">
<a href="#" onclick="swapDivClassname(this, 'highlightTaxonNameWrapper', 'highlightTaxonNameWrapper')">
Taxon names
</a>
</td>
<td class="citationsButton">
<a href="#" onclick="swapDivClassname(this, 'highlightCitationWrapper', 'highlightCitationWrapper')">
Citations
</a>
</td>
<td class="offButton">
<a class="inactive" href="#" onclick="swapDivClassname(this, 'highlightWrapper', 'highlightWrapper')">
Turn highlighting On/Off
</a>
</td>
</tr>
</tbody>
</table>
<div class="linksButtonWrapper">
<div class="pathTopLeftScrollBtn">
<a class="LeftScrollBtnLinkInactive" id="scrollLeftBtn" onmouseout="stopScroll()" onmouseover="startScroll(1)">
</a>
</div>
<div class="pathScrollHolder" id="scrollHolderDiv" style="width: 100%;">
<div class="pathScrollDiv" id="scrollDiv" style="width: 100%; left: 0px;">
<table cellpadding="3" cellspacing="0" class="linksButtonHolder" id="linksButtonHolder">
<tbody>
<tr>
<td class="linksButtonHolder">
<a href="#top">
Top
</a>
</td>
<td class="linksButtonHolder">
<a href="#abstract" onclick="">
Abstract
</a>
</td>
<td class="linksButtonHolder">
<a href="#introduction" onclick="">
Introduction
</a>
</td>
<td class="linksButtonHolder">
<a href="#materialandmethods" onclick="">
Material and methods
</a>
</td>
<td class="linksButtonHolder">
<a href="#speciestreatment" onclick="">
Species treatment
</a>
</td>
<td class="linksButtonHolder">
<a href="#identificationkey" onclick="">
Identification key
</a>
</td>
<td class="linksButtonHolder">
<a href="#phylogeneticanalysisofmoleculardata" onclick="">
Phylogenetic analysis of molecular data
</a>
</td>
<td class="linksButtonHolder">
<a href="#discussion" onclick="">
Discussion
</a>
</td>
<td class="linksButtonHolder">
<a href="#acknowledgements" onclick="">
Acknowledgements
</a>
</td>
<td class="linksButtonHolder">
<a href="#references" onclick="">
References
</a>
</td>
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<div class="doiNumber">
ZooKeys 103: 49–62, doi: 10.3897/ZooKeys.103.1124
</div>
<div class="mainTitle">
A new endogean, anophthalmous species of
<i>
Parazuphium
</i>
Jeannel from Northern Morocco (Coleoptera, Carabidae), with
new molecular data for the tribe Zuphiini
</div>
<div class="authors">
Carmelo Andújar
<span class="specialSymbols">
1,†
</span>
, Carles Hernando
<span class="specialSymbols">
2,‡
</span>
, Ignacio Ribera
<span class="specialSymbols">
3,§
</span>
</div>
<div class="affilations">
<div class="affiliationRow">
1
<i>
Departamento de Zoología y Antropología Física. Facultad de Veterinaria, Universidad de Murcia. 30071 Murcia, Spain
</i>
</div>
<div class="affiliationRow">
2
<i>
Apartado de Correos 118, E-08911 Badalona, Barcelona, Spain
</i>
</div>
<div class="affiliationRow">
3
<i>
Institute of Evolutionary Biology (CSIC-UPF), Passeig Maritim de la Barceloneta 37-49, 08003 Barcelona, Spain
</i>
</div>
</div>
<div class="uris">
<div class="uriRow">
†
<a href="http://zoobank.org/?lsid=urn:lsid:zoobank.org:author:6F2F5C16-AF3E-40C6-9C70-40D51BF0FA66" target="_blank">
urn:lsid:zoobank.org:author:6F2F5C16-AF3E-40C6-9C70-40D51BF0FA66
</a>
</div>
<div class="uriRow">
‡
<a href="http://zoobank.org/?lsid=urn:lsid:zoobank.org:author:12EC1600-4D8F-471E-B3C6-A72000BCC1B8" target="_blank">
urn:lsid:zoobank.org:author:12EC1600-4D8F-471E-B3C6-A72000BCC1B8
</a>
</div>
<div class="uriRow">
§
<a href="http://zoobank.org/?lsid=urn:lsid:zoobank.org:author:8CF31E58-DC2A-4F16-9B0B-616FAF93AD0D" target="_blank">
urn:lsid:zoobank.org:author:8CF31E58-DC2A-4F16-9B0B-616FAF93AD0D
</a>
</div>
</div>
<div class="authorNotes">
<div class="authorNotesRow">
<p>
Corresponding author: Carmelo Andújar (
<a href="mailto:candujar@um.es">
candujar@um.es
</a>
).
</p>
<p>
Academic editor: Terry Erwin
</p>
</div>
</div>
<div class="lMetaDates">
<span class="metaDate">
<span class="dateLabel">
received
</span>
14 February 2011
</span>
<span class="dateSeparator">
|
</span>
<span class="metaDate">
<span class="dateLabel">
accepted
</span>
28 April 2011
</span>
<span class="dateSeparator">
|
</span>
<span class="metaDate">
<span class="dateLabel">
Published
</span>
10 June 2011
</span>
<p>
</p>
<br/>
<div class="rifgtsrow">
<p>
(C) 2011 Carmelo Andújar. This is an open access article distributed under the terms of the
<a href="http://creativecommons.org/licenses/by/3.0/" target="_blank">
Creative Commons Attribution License
</a>
,
which permits unrestricted use, distribution, and reproduction in any
medium, provided the original author and source are credited.
</p>
<br/>
<p>
For reference, use of the paginated PDF or printed version of this article is recommended.
</p>
</div>
</div>
<a name="abstract">
</a>
<div class="abstractHolder">
<div class="abstractText">
<span class="node_label">
Abstract
</span>
<p>
A new species of the genus
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
</span>
</span>
</i>
(
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Coleoptera" onmouseout="hideBaloonEvent('Coleoptera', event)" onmouseover="showBaloon(2, 'Coleoptera', event)" target="_blank">
Coleoptera
</a>
</span>
</span>
,
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Carabidae" onmouseout="hideBaloonEvent('Carabidae', event)" onmouseover="showBaloon(2, 'Carabidae', event)" target="_blank">
Carabidae
</a>
</span>
</span>
,
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Zuphiini" onmouseout="hideBaloonEvent('Zuphiini', event)" onmouseover="showBaloon(2, 'Zuphiini', event)" target="_blank">
Zuphiini
</a>
</span>
</span>
),
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" onmouseout="hideBaloonEvent('Parazuphium_aguilerai', event)" onmouseover="showBaloon(2, 'Parazuphium_aguilerai', event)" target="_blank">
aguilerai
</a>
</span>
</span>
</i>
<b>
sp. n.
</b>
,
is described from the Tingitan peninsula in North Morocco. The only
known specimen was found under a large deeply buried boulder, and
belongs to an anophthalmous, depigmented and flattened species. This is
the second species of blind
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
</span>
</span>
</i>
known so far, the other being
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20feloi" onmouseout="hideBaloonEvent('Parazuphium_feloi', event)" onmouseover="showBaloon(2, 'Parazuphium_feloi', event)" target="_blank">
feloi
</a>
</span>
</span>
</i>
Machado 1998 from a lava tube in the Canary Islands. Molecular data of the only known
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" onmouseout="hideBaloonEvent('Parazuphium_aguilerai', event)" onmouseover="showBaloon(2, 'Parazuphium_aguilerai', event)" target="_blank">
aguilerai
</a>
</span>
</span>
</i>
sp. n. specimen are provided, and a reconstructed phylogeny based on these molecular data confirms its inclusion inside
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Zuphiini" onmouseout="hideBaloonEvent('Zuphiini', event)" onmouseover="showBaloon(2, 'Zuphiini', event)" target="_blank">
Zuphiini
</a>
</span>
</span>
within
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Harpalinae" onmouseout="hideBaloonEvent('Harpalinae', event)" onmouseover="showBaloon(2, 'Harpalinae', event)" target="_blank">
Harpalinae
</a>
</span>
</span>
. Identification keys to the Mediterranean and Macaronesian species of
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
</span>
</span>
</i>
are provided.
</p>
</div>
</div>
<div class="keywordsHolder">
<div class="keywordsText">
<span class="node_label">
Keywords
</span>
<p>
Taxonomy, new species,
<i>
Parazuphium
</i>
, identification key, endogean fauna, molecular phylogeny
</p>
</div>
</div>
<div class=" secHolder">
<a name="introduction">
</a>
<span class="node_title">
Introduction
</span>
<p>
The genus
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
</span>
</span>
</i>
Latreille (
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Coleoptera" onmouseout="hideBaloonEvent('Coleoptera', event)" onmouseover="showBaloon(2, 'Coleoptera', event)" target="_blank">
Coleoptera
</a>
</span>
</span>
,
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Dryptinae" onmouseout="hideBaloonEvent('Dryptinae', event)" onmouseover="showBaloon(2, 'Dryptinae', event)" target="_blank">
Dryptinae
</a>
</span>
</span>
,
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Zuphiini" onmouseout="hideBaloonEvent('Zuphiini', event)" onmouseover="showBaloon(2, 'Zuphiini', event)" target="_blank">
Zuphiini
</a>
</span>
</span>
)
is characterized by the presence in the aedeagus of a strong ventral
constriction between the basal and the distal part of the median lobe, a
unique structure within the
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Carabidae" onmouseout="hideBaloonEvent('Carabidae', event)" onmouseover="showBaloon(2, 'Carabidae', event)" target="_blank">
Carabidae
</a>
</span>
</span>
(
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B1')" onmouseover="showBaloon(1, 'B1', event)">
Antoine 1962
</span>
,
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B13')" onmouseover="showBaloon(1, 'B13', event)">
Mateu 1993
</span>
). The genus currently includes ca. 40 species (
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B9')" onmouseover="showBaloon(1, 'B9', event)">
Lorenz 2005
</span>
) in the Old World and Australia (
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B13')" onmouseover="showBaloon(1, 'B13', event)">
Mateu 1993
</span>
), arranged in three subgenera: 1)
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Neozuphium" onmouseout="hideBaloonEvent('Neozuphium', event)" onmouseover="showBaloon(2, 'Neozuphium', event)" target="_blank">
Neozuphium
</a>
</span>
</span>
</i>
Hürka, with a single species,
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Neozuphium" onmouseout="hideBaloonEvent('Neozuphium', event)" onmouseover="showBaloon(2, 'Neozuphium', event)" target="_blank">
(Neozuphium)
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20%28Neozuphium%29%20damascenum" onmouseout="hideBaloonEvent('Parazuphium_(Neozuphium)_damascenum', event)" onmouseover="showBaloon(2, 'Parazuphium_(Neozuphium)_damascenum', event)" target="_blank">
damascenum
</a>
</span>
</span>
</i>
(Fairmaire) (
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B12')" onmouseover="showBaloon(1, 'B12', event)">
Mateu 1988
</span>
), with a wide Palaearctic distribution from Central Asia and the Mediterranean to the Canaries (
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B5')" onmouseover="showBaloon(1, 'B5', event)">
Hürka 1982
</span>
,
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B12')" onmouseover="showBaloon(1, 'B12', event)">
Mateu 1988
</span>
,
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B10')" onmouseover="showBaloon(1, 'B10', event)">
Machado 1992
</span>
,
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B7')" onmouseover="showBaloon(1, 'B7', event)">
Lencina and Serrano 1995
</span>
); 2)
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Austrozuphium" onmouseout="hideBaloonEvent('Austrozuphium', event)" onmouseover="showBaloon(2, 'Austrozuphium', event)" target="_blank">
Austrozuphium
</a>
</span>
</span>
</i>
Baehr, with five Australasian species (
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B2')" onmouseover="showBaloon(1, 'B2', event)">
Baehr 1985
</span>
) and 3)
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
</span>
</span>
</i>
s.str. Jeannel, with 32 species distributed through the Old World (
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B13')" onmouseover="showBaloon(1, 'B13', event)">
Mateu 1993
</span>
). The recent catalogues of
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B8')" onmouseover="showBaloon(1, 'B8', event)">
Löbl and Smetana (2003)
</span>
and
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B9')" onmouseover="showBaloon(1, 'B9', event)">
Lorenz (2005)
</span>
do not include the synonymies of
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Neozuphium" onmouseout="hideBaloonEvent('Neozuphium', event)" onmouseover="showBaloon(2, 'Neozuphium', event)" target="_blank">
(Neozuphium)
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20%28Neozuphium%29%20bactrianum" onmouseout="hideBaloonEvent('Parazuphium_(Neozuphium)_bactrianum', event)" onmouseover="showBaloon(2, 'Parazuphium_(Neozuphium)_bactrianum', event)" target="_blank">
bactrianum
</a>
</span>
</span>
</i>
(K. Daniel & F. Daniel) and
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Neozuphium" onmouseout="hideBaloonEvent('Neozuphium', event)" onmouseover="showBaloon(2, 'Neozuphium', event)" target="_blank">
(Neozuphium)
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20%28Neozuphium%29%20novaki" onmouseout="hideBaloonEvent('Parazuphium_(Neozuphium)_novaki', event)" onmouseover="showBaloon(2, 'Parazuphium_(Neozuphium)_novaki', event)" target="_blank">
novaki
</a>
</span>
</span>
</i>
(G. Müller) with
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20damascenum" onmouseout="hideBaloonEvent('Parazuphium_damascenum', event)" onmouseover="showBaloon(2, 'Parazuphium_damascenum', event)" target="_blank">
damascenum
</a>
</span>
</span>
</i>
(Mateu [1988]).
</p>
<p>
The species of the genus seem to be associated with deep soil or the soil crevices near rivers or temporary flooded areas (
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B2')" onmouseover="showBaloon(1, 'B2', event)">
Baehr 1985
</span>
,
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B10')" onmouseover="showBaloon(1, 'B10', event)">
Machado 1992
</span>
),
and generally show a flattened habitus, some degree of depigmentation
and microphthalmy. Some species are known from caves, one of them
being the only previously known blind species of the genus (
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20feloi" onmouseout="hideBaloonEvent('Parazuphium_feloi', event)" onmouseover="showBaloon(2, 'Parazuphium_feloi', event)" target="_blank">
feloi
</a>
</span>
</span>
</i>
Machado, from the Canary islands) (
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B11')" onmouseover="showBaloon(1, 'B11', event)">
Machado 1998
</span>
).
</p>
<p>
During an entomological expedition to North Morocco we found the single specimen of a new species of
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
</span>
</span>
</i>
,
anophthalmous and with strong modifications apparently related to its
endogean habitat. Despite an attempt to collect additional material the
following year no other specimen was found, possibly due to the
endogean habits of this species. We describe the species here, and
provide some molecular data to characterize it and to postulate its
phylogenetic position among the
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Zuphiini" onmouseout="hideBaloonEvent('Zuphiini', event)" onmouseover="showBaloon(2, 'Zuphiini', event)" target="_blank">
Zuphiini
</a>
</span>
</span>
for which genetic data are available (
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B17')" onmouseover="showBaloon(1, 'B17', event)">
Ribera et al. 2006
</span>
).
</p>
<a name="materialandmethods">
</a>
<span class="node_title">
Material and methods
</span>
<p>
The unique specimen was killed and stored in absolute
ethanol in the field, and total DNA was extracted using the QIAGEN
Dneasy tissue kit (Qiagen, Hilden, Germany), without destroying the
external cuticle. The extracted specimen was mounted in DMHF (Dimethyl
Hydantoin-Formaldehyde) on a transparent acetate label. For the
morphological study and photographs we used a Zeiss Stemi 2000C
Trinocular Zoom Stereomicroscope with Spot Insight Firewire digital
camera and software.
</p>
<a name="molecularmethods">
</a>
<span class="node_title1">
Molecular methods
</span>
<p>
Total genomic DNA for the single specimen of
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" onmouseout="hideBaloonEvent('Parazuphium_aguilerai', event)" onmouseover="showBaloon(2, 'Parazuphium_aguilerai', event)" target="_blank">
aguilerai
</a>
</span>
</span>
</i>
sp. n. was extracted using QIAGEN Dneasy tissue kit (Qiagen, Hilden,
Germany). To characterize the new species we amplified fragments of six
genes, four mitochondrial and two nuclear: 3' end of cytochrome c
oxidase subunit (
<i>
cox1
</i>
); a single fragment including the 3' end of the large ribosomal unit (
<i>
rrnL
</i>
), the whole tRNA-Leu gene (
<i>
trnL
</i>
) and the 5' end of the NADH dehydrogenase 1 (
<i>
nad1
</i>
); 5' end of the small ribosomal unit, 18S rRNA (
<i>
SSU
</i>
); and an internal fragment of the large ribosomal unit, 28S rRNA (
<i>
LSU
</i>
). Primers used are given in
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('T1')" onmouseover="showBaloon(4, 'T1', event)">
Table 1
</span>
. Additionally, we extracted DNA from one specimen of
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
</span>
</span>
</i>
cf.
<i>
baeticum
</i>
(K. and J. Daniel 1898),
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Zuphium" onmouseout="hideBaloonEvent('Zuphium', event)" onmouseover="showBaloon(2, 'Zuphium', event)" target="_blank">
Zuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Zuphium%20olens" onmouseout="hideBaloonEvent('Zuphium_olens', event)" onmouseover="showBaloon(2, 'Zuphium_olens', event)" target="_blank">
olens
</a>
</span>
</span>
</i>
Rossi 1790,
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Ildobates" onmouseout="hideBaloonEvent('Ildobates', event)" onmouseover="showBaloon(2, 'Ildobates', event)" target="_blank">
Ildobates
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Ildobates%20neboti" onmouseout="hideBaloonEvent('Ildobates_neboti', event)" onmouseover="showBaloon(2, 'Ildobates_neboti', event)" target="_blank">
neboti
</a>
</span>
</span>
</i>
Español 1966 and several other outgroups among
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Carabidae" onmouseout="hideBaloonEvent('Carabidae', event)" onmouseover="showBaloon(2, 'Carabidae', event)" target="_blank">
Carabidae
</a>
</span>
</span>
(
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('T2')" onmouseover="showBaloon(4, 'T2', event)">
Table 2
</span>
),
which were amplified for the same molecular gene fragments. PCR
reactions were made using PuReTaq Ready-To-Go PCR beads (GE Healthcare,
UK) and standard conditions [39 cycles using 48–50°C as annealing
temperature]. New sequences have been deposited in GenBank (NCBI) with
Acc. Nos JF778779-JF778845. Each individual gene matrix was aligned in
MAFFT with the Q-ins-i option and default parameters. The four genes
fragments were concatenated to get a final dataset of 20 taxa and 3376
bp that was employed in phylogenetic analyses.
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('T2')" onmouseover="showBaloon(4, 'T2', event)">
Table 2
</span>
shows taxa information, source and accession number for each DNA sequence.
</p>
<div class="tableWrapHolder Table">
<span class="node_label">
Table 1.
</span>
<p>
Primers used in the study. F, forward; R, reverse. Length refers to the aligned matrix.
</p>
<div class="tableDiv" id="table_T1">
<table class="innerTable Table">
<tbody>
<tr>
<th class="cell_1" colspan="1">
Type DNA
</th>
<th class="cell_2" colspan="1">
Gene
</th>
<th class="cell_3" colspan="1">
Length
</th>
<th class="cell_4" colspan="1">
Primer
</th>
<th class="cell_5" colspan="1">
S
</th>
<th class="cell_6" colspan="1">
Primer sequence (5'- 3')
</th>
<th class="cell_7" colspan="1">
Described in:
</th>
</tr>
<tr>
<td class="cell_1" colspan="1">
Mitochondrial protein coding
</td>
<td class="cell_2" colspan="1">
cox1
</td>
<td class="cell_3" colspan="1">
755
</td>
<td class="cell_4" colspan="1">
Jerry (M202)
</td>
<td class="cell_5" colspan="1">
F
</td>
<td class="cell_6" colspan="1">
CAACATTTATTT-TGATTTTTTGG
</td>
<td class="cell_7" colspan="1">
(
<span class="xrefLink node_citation">
Simon et al. 1994
</span>
)
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
Pat (M70)
</td>
<td class="cell_2" colspan="1">
R
</td>
<td class="cell_3" colspan="1">
TCCA(A)TGCACTA-ATCTGCCATATTA
</td>
<td class="cell_4" colspan="1">
(
<span class="xrefLink node_citation">
Simon et al. 1994
</span>
)
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
Mitochondrial ribosomal
</td>
<td class="cell_2" colspan="1">
rrnL
</td>
<td class="cell_3" colspan="1">
744
</td>
<td class="cell_4" colspan="1">
16SaR (M14)
</td>
<td class="cell_5" colspan="1">
F
</td>
<td class="cell_6" colspan="1">
CGCCTGTTTA-WCAAAAACAT
</td>
<td class="cell_7" colspan="1">
(
<span class="xrefLink node_citation">
Simon et al. 1994
</span>
)
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
16s-ND1a (M223)
</td>
<td class="cell_2" colspan="1">
R
</td>
<td class="cell_3" colspan="1">
GGTCCCTTACGAA-TTTGAATATATCCT
</td>
<td class="cell_4" colspan="1">
(
<span class="xrefLink node_citation">
Simon et al. 1994
</span>
)
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
Nuclear ribosomal
</td>
<td class="cell_2" colspan="1">
LSU
</td>
<td class="cell_3" colspan="1">
1240
</td>
<td class="cell_4" colspan="1">
LS58F (D1)
</td>
<td class="cell_5" colspan="1">
F
</td>
<td class="cell_6" colspan="1">
GGGAGGAAA-AGAAACTAAC
</td>
<td class="cell_7" colspan="1">
(
<span class="xrefLink node_citation">
Ober 2002
</span>
)
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
LS998R (D3)
</td>
<td class="cell_2" colspan="1">
R
</td>
<td class="cell_3" colspan="1">
GCATAGTTC-ACCATCTTTC
</td>
<td class="cell_4" colspan="1">
(
<span class="xrefLink node_citation">
Ober 2002
</span>
)
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
Nuclear ribosomal
</td>
<td class="cell_2" colspan="1">
SSU
</td>
<td class="cell_3" colspan="1">
625
</td>
<td class="cell_4" colspan="1">
5'b5.0
</td>
<td class="cell_5" colspan="1">
F
</td>
<td class="cell_6" colspan="1">
GACAACCTGGTT-GATCCTGCCAGT
</td>
<td class="cell_7" colspan="1">
(
<span class="xrefLink node_citation">
Shull et al. 2001
</span>
)
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
R
</td>
<td class="cell_2" colspan="1">
TAACCGCAA-CAACTTTAAT
</td>
<td class="cell_3" colspan="1">
(
<span class="xrefLink node_citation">
Shull et al. 2001
</span>
)
</td>
</tr>
</tbody>
</table>
</div>
<table cellpadding="0" cellspacing="0" class="innerTable Table">
<tbody>
<tr>
<th class="cell_1" colspan="1">
Type DNA
</th>
<th class="cell_2" colspan="1">
Gene
</th>
<th class="cell_3" colspan="1">
Length
</th>
<th class="cell_4" colspan="1">
Primer
</th>
<th class="cell_5" colspan="1">
S
</th>
<th class="cell_6" colspan="1">
Primer sequence (5'- 3')
</th>
<th class="cell_7" colspan="1">
Described in:
</th>
</tr>
<tr>
<td class="cell_1" colspan="1">
Mitochondrial protein coding
</td>
<td class="cell_2" colspan="1">
cox1
</td>
<td class="cell_3" colspan="1">
755
</td>
<td class="cell_4" colspan="1">
Jerry (M202)
</td>
<td class="cell_5" colspan="1">
F
</td>
<td class="cell_6" colspan="1">
CAACATTTATTT-TGATTTTTTGG
</td>
<td class="cell_7" colspan="1">
(
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B21')" onmouseover="showBaloon(1, 'B21', event)">
Simon et al. 1994
</span>
)
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
Pat (M70)
</td>
<td class="cell_2" colspan="1">
R
</td>
<td class="cell_3" colspan="1">
TCCA(A)TGCACTA-ATCTGCCATATTA
</td>
<td class="cell_4" colspan="1">
(
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B21')" onmouseover="showBaloon(1, 'B21', event)">
Simon et al. 1994
</span>
)
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
Mitochondrial ribosomal
</td>
<td class="cell_2" colspan="1">
rrnL
</td>
<td class="cell_3" colspan="1">
744
</td>
<td class="cell_4" colspan="1">
16SaR (M14)
</td>
<td class="cell_5" colspan="1">
F
</td>
<td class="cell_6" colspan="1">
CGCCTGTTTA-WCAAAAACAT
</td>
<td class="cell_7" colspan="1">
(
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B21')" onmouseover="showBaloon(1, 'B21', event)">
Simon et al. 1994
</span>
)
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
16s-ND1a (M223)
</td>
<td class="cell_2" colspan="1">
R
</td>
<td class="cell_3" colspan="1">
GGTCCCTTACGAA-TTTGAATATATCCT
</td>
<td class="cell_4" colspan="1">
(
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B21')" onmouseover="showBaloon(1, 'B21', event)">
Simon et al. 1994
</span>
)
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
Nuclear ribosomal
</td>
<td class="cell_2" colspan="1">
LSU
</td>
<td class="cell_3" colspan="1">
1240
</td>
<td class="cell_4" colspan="1">
LS58F (D1)
</td>
<td class="cell_5" colspan="1">
F
</td>
<td class="cell_6" colspan="1">
GGGAGGAAA-AGAAACTAAC
</td>
<td class="cell_7" colspan="1">
(
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B14')" onmouseover="showBaloon(1, 'B14', event)">
Ober 2002
</span>
)
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
LS998R (D3)
</td>
<td class="cell_2" colspan="1">
R
</td>
<td class="cell_3" colspan="1">
GCATAGTTC-ACCATCTTTC
</td>
<td class="cell_4" colspan="1">
(
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B14')" onmouseover="showBaloon(1, 'B14', event)">
Ober 2002
</span>
)
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
Nuclear ribosomal
</td>
<td class="cell_2" colspan="1">
SSU
</td>
<td class="cell_3" colspan="1">
625
</td>
<td class="cell_4" colspan="1">
5'b5.0
</td>
<td class="cell_5" colspan="1">
F
</td>
<td class="cell_6" colspan="1">
GACAACCTGGTT-GATCCTGCCAGT
</td>
<td class="cell_7" colspan="1">
(
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B20')" onmouseover="showBaloon(1, 'B20', event)">
Shull et al. 2001
</span>
)
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
R
</td>
<td class="cell_2" colspan="1">
TAACCGCAA-CAACTTTAAT
</td>
<td class="cell_3" colspan="1">
(
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B20')" onmouseover="showBaloon(1, 'B20', event)">
Shull et al. 2001
</span>
)
</td>
</tr>
</tbody>
</table>
</div>
<div class="tableWrapHolder Table">
<span class="node_label">
Table 2.
</span>
<p>
Species, locality of collection, voucher reference and accession numbers for each sequence.
</p>
<div class="tableDiv" id="table_T2">
<table class="innerTable Table">
<tbody>
<tr>
<th class="cell_1" colspan="1">
Especie
</th>
<th class="cell_2" colspan="1">
Locality
</th>
<th class="cell_3" colspan="1">
Voucher
</th>
<th class="cell_4" colspan="1">
cox1
</th>
<th class="cell_5" colspan="1">
rrnL
</th>
<th class="cell_6" colspan="1">
LSU
</th>
<th class="cell_7" colspan="1">
SSU
</th>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Laemostenus" target="_blank">
Laemostenus
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Laemostenus%20terricola" target="_blank">
terricola
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Alicante, Spain
</td>
<td class="cell_3" colspan="1">
1583BG
</td>
<td class="cell_4" colspan="1">
JF778779
</td>
<td class="cell_5" colspan="1">
JF778796
</td>
<td class="cell_6" colspan="1">
JF778812
</td>
<td class="cell_7" colspan="1">
JF778829
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Leistus" target="_blank">
Leistus
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Leistus%20spinibarbis" target="_blank">
spinibarbis
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Albacete, Spain
</td>
<td class="cell_3" colspan="1">
1581BG
</td>
<td class="cell_4" colspan="1">
JF778780
</td>
<td class="cell_5" colspan="1">
JF778797
</td>
<td class="cell_6" colspan="1">
JF778813
</td>
<td class="cell_7" colspan="1">
JF778830
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Calosoma" target="_blank">
Calosoma
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Calosoma%20sycophanta" target="_blank">
sycophanta
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Albacete, Spain
</td>
<td class="cell_3" colspan="1">
1590BG
</td>
<td class="cell_4" colspan="1">
JF778781
</td>
<td class="cell_5" colspan="1">
JF778798
</td>
<td class="cell_6" colspan="1">
JF778814
</td>
<td class="cell_7" colspan="1">
JF778831
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Carabus" target="_blank">
Carabus
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Eucarabus" target="_blank">
(Eucarabus)
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Carabus%20%28Eucarabus%29%20deyrollei" target="_blank">
deyrollei
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Lugo, Spain
</td>
<td class="cell_3" colspan="1">
1553BG
</td>
<td class="cell_4" colspan="1">
JF778782
</td>
<td class="cell_5" colspan="1">
JF778799
</td>
<td class="cell_6" colspan="1">
JF778815
</td>
<td class="cell_7" colspan="1">
JF778832
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Carabus" target="_blank">
Carabus
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Limnocarabaus" target="_blank">
(Limnocarabaus)
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Carabus%20%28Limnocarabaus%29%20clathratus" target="_blank">
clathratus
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Susuz, Turkey
</td>
<td class="cell_3" colspan="1">
1600BG
</td>
<td class="cell_4" colspan="1">
JF778783
</td>
<td class="cell_5" colspan="1">
JF778800
</td>
<td class="cell_6" colspan="1">
JF778816
</td>
<td class="cell_7" colspan="1">
JF778833
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Dixus" target="_blank">
Dixus
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Dixus%20capito" target="_blank">
capito
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Albacete, Spain
</td>
<td class="cell_3" colspan="1">
1578BG
</td>
<td class="cell_4" colspan="1">
JF778784
</td>
<td class="cell_5" colspan="1">
N/A
</td>
<td class="cell_6" colspan="1">
JF778817
</td>
<td class="cell_7" colspan="1">
JF778834
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Pseudotrechus" target="_blank">
Pseudotrechus
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Pseudotrechus%20mutilatus" target="_blank">
mutilatus
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Cádiz, Spain
</td>
<td class="cell_3" colspan="1">
36_EN
</td>
<td class="cell_4" colspan="1">
JF778785
</td>
<td class="cell_5" colspan="1">
JF778801
</td>
<td class="cell_6" colspan="1">
JF778818
</td>
<td class="cell_7" colspan="1">
JF778835
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Licinus" target="_blank">
Licinus
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Licinus%20punctatulus" target="_blank">
punctatulus
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Alicante, Spain
</td>
<td class="cell_3" colspan="1">
1582BG
</td>
<td class="cell_4" colspan="1">
JF778786
</td>
<td class="cell_5" colspan="1">
JF778802
</td>
<td class="cell_6" colspan="1">
JF778819
</td>
<td class="cell_7" colspan="1">
JF778836
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Elaphropus" target="_blank">
Elaphropus
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Tachyura" target="_blank">
(Tachyura)
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Elaphropus%20%28Tachyura%29%20parvulus" target="_blank">
parvulus
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Pays Zaer Zaine, Morocco
</td>
<td class="cell_3" colspan="1">
64_EN
</td>
<td class="cell_4" colspan="1">
JF778787
</td>
<td class="cell_5" colspan="1">
JF778803
</td>
<td class="cell_6" colspan="1">
JF778820
</td>
<td class="cell_7" colspan="1">
JF778837
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Bembidion" target="_blank">
Bembidion
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Peryphus" target="_blank">
(Peryphus)
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Bembidion%20%28Peryphus%29%20hispanicum" target="_blank">
hispanicum
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Pays Zaer Zaine, Morocco
</td>
<td class="cell_3" colspan="1">
62_EN
</td>
<td class="cell_4" colspan="1">
N/A
</td>
<td class="cell_5" colspan="1">
JF778804
</td>
<td class="cell_6" colspan="1">
JF778821
</td>
<td class="cell_7" colspan="1">
JF778838
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Bembidion" target="_blank">
Bembidion
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Emphanes" target="_blank">
(Emphanes)
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Bembidion%20%28Emphanes%29%20latiplaga" target="_blank">
latiplaga
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Pays Zaer Zaine, Morocco
</td>
<td class="cell_3" colspan="1">
65_EN
</td>
<td class="cell_4" colspan="1">
JF778788
</td>
<td class="cell_5" colspan="1">
N/A
</td>
<td class="cell_6" colspan="1">
JF778822
</td>
<td class="cell_7" colspan="1">
JF778839
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Perileptus" target="_blank">
Perileptus
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Perileptus%20aerolatus" target="_blank">
aerolatus
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Agadir, Morocco
</td>
<td class="cell_3" colspan="1">
MNHN-AF113
</td>
<td class="cell_4" colspan="1">
GQ293688
</td>
<td class="cell_5" colspan="1">
FR729593
</td>
<td class="cell_6" colspan="1">
GQ293625
</td>
<td class="cell_7" colspan="1">
GQ293503
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Trechus" target="_blank">
Trechus
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Trechus%20quadristriatus" target="_blank">
quadristriatus
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Huesca, Spain
</td>
<td class="cell_3" colspan="1">
MNHN-AF96
</td>
<td class="cell_4" colspan="1">
FR733908
</td>
<td class="cell_5" colspan="1">
GQ293743
</td>
<td class="cell_6" colspan="1">
GQ293619
</td>
<td class="cell_7" colspan="1">
GQ293534
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Typhloreicheia" target="_blank">
Typhloreicheia
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Typhloreicheia%20laurentii" target="_blank">
laurentii
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Sardinia, Italy
</td>
<td class="cell_3" colspan="1">
56_EN
</td>
<td class="cell_4" colspan="1">
JF778789
</td>
<td class="cell_5" colspan="1">
JF778805
</td>
<td class="cell_6" colspan="1">
JF778823
</td>
<td class="cell_7" colspan="1">
JF778840
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Dyschiriodes" target="_blank">
Dyschiriodes
</a>
</span>
</span>
</i>
sp.
</td>
<td class="cell_2" colspan="1">
Pays Zaer Zaine, Morocco
</td>
<td class="cell_3" colspan="1">
63_EN
</td>
<td class="cell_4" colspan="1">
JF778790
</td>
<td class="cell_5" colspan="1">
JF778806
</td>
<td class="cell_6" colspan="1">
JF778824
</td>
<td class="cell_7" colspan="1">
JF778841
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Nebria" target="_blank">
Nebria
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Nebria%20salina" target="_blank">
salina
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Albacete, Spain
</td>
<td class="cell_3" colspan="1">
1579BG
</td>
<td class="cell_4" colspan="1">
JF778791
</td>
<td class="cell_5" colspan="1">
JF778807
</td>
<td class="cell_6" colspan="1">
JF778825
</td>
<td class="cell_7" colspan="1">
JF778842
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Ildobates" target="_blank">
Ildobates
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Ildobates%20neboti" target="_blank">
neboti
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Castellón, Spain
</td>
<td class="cell_3" colspan="1">
MNCN-6409
</td>
<td class="cell_4" colspan="1">
JF778792
</td>
<td class="cell_5" colspan="1">
JF778808
</td>
<td class="cell_6" colspan="1">
AM051084
</td>
<td class="cell_7" colspan="1">
DQ130051
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Drypta" target="_blank">
Drypta
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Drypta%20dentata" target="_blank">
dentata
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Ciudad Real, Spain
</td>
<td class="cell_3" colspan="1">
98_EN
</td>
<td class="cell_4" colspan="1">
N/A
</td>
<td class="cell_5" colspan="1">
JF778809
</td>
<td class="cell_6" colspan="1">
N/A
</td>
<td class="cell_7" colspan="1">
N/A
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Zuphium" target="_blank">
Zuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Zuphium%20olens" target="_blank">
olens
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Murcia, Spain
</td>
<td class="cell_3" colspan="1">
97_EN
</td>
<td class="cell_4" colspan="1">
JF778793
</td>
<td class="cell_5" colspan="1">
N/A
</td>
<td class="cell_6" colspan="1">
JF778826
</td>
<td class="cell_7" colspan="1">
JF778843
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20cf.%20baeticum" target="_blank">
cf.
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20cf.%20baeticum" target="_blank">
baeticum
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Castellón, Spain
</td>
<td class="cell_3" colspan="1">
87_EN
</td>
<td class="cell_4" colspan="1">
JF778794
</td>
<td class="cell_5" colspan="1">
JF778810
</td>
<td class="cell_6" colspan="1">
JF778827
</td>
<td class="cell_7" colspan="1">
JF778844
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" target="_blank">
aguilerai
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Tanger, Morocco
</td>
<td class="cell_3" colspan="1">
31_EN
</td>
<td class="cell_4" colspan="1">
JF778795
</td>
<td class="cell_5" colspan="1">
JF778811
</td>
<td class="cell_6" colspan="1">
JF778828
</td>
<td class="cell_7" colspan="1">
JF778845
</td>
</tr>
</tbody>
</table>
</div>
<table cellpadding="0" cellspacing="0" class="innerTable Table">
<tbody>
<tr>
<th class="cell_1" colspan="1">
Especie
</th>
<th class="cell_2" colspan="1">
Locality
</th>
<th class="cell_3" colspan="1">
Voucher
</th>
<th class="cell_4" colspan="1">
cox1
</th>
<th class="cell_5" colspan="1">
rrnL
</th>
<th class="cell_6" colspan="1">
LSU
</th>
<th class="cell_7" colspan="1">
SSU
</th>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Laemostenus" onmouseout="hideBaloonEvent('Laemostenus', event)" onmouseover="showBaloon(2, 'Laemostenus', event)" target="_blank">
Laemostenus
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Laemostenus%20terricola" onmouseout="hideBaloonEvent('Laemostenus_terricola', event)" onmouseover="showBaloon(2, 'Laemostenus_terricola', event)" target="_blank">
terricola
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Alicante, Spain
</td>
<td class="cell_3" colspan="1">
1583BG
</td>
<td class="cell_4" colspan="1">
JF778779
</td>
<td class="cell_5" colspan="1">
JF778796
</td>
<td class="cell_6" colspan="1">
JF778812
</td>
<td class="cell_7" colspan="1">
JF778829
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Leistus" onmouseout="hideBaloonEvent('Leistus', event)" onmouseover="showBaloon(2, 'Leistus', event)" target="_blank">
Leistus
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Leistus%20spinibarbis" onmouseout="hideBaloonEvent('Leistus_spinibarbis', event)" onmouseover="showBaloon(2, 'Leistus_spinibarbis', event)" target="_blank">
spinibarbis
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Albacete, Spain
</td>
<td class="cell_3" colspan="1">
1581BG
</td>
<td class="cell_4" colspan="1">
JF778780
</td>
<td class="cell_5" colspan="1">
JF778797
</td>
<td class="cell_6" colspan="1">
JF778813
</td>
<td class="cell_7" colspan="1">
JF778830
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Calosoma" onmouseout="hideBaloonEvent('Calosoma', event)" onmouseover="showBaloon(2, 'Calosoma', event)" target="_blank">
Calosoma
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Calosoma%20sycophanta" onmouseout="hideBaloonEvent('Calosoma_sycophanta', event)" onmouseover="showBaloon(2, 'Calosoma_sycophanta', event)" target="_blank">
sycophanta
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Albacete, Spain
</td>
<td class="cell_3" colspan="1">
1590BG
</td>
<td class="cell_4" colspan="1">
JF778781
</td>
<td class="cell_5" colspan="1">
JF778798
</td>
<td class="cell_6" colspan="1">
JF778814
</td>
<td class="cell_7" colspan="1">
JF778831
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Carabus" onmouseout="hideBaloonEvent('Carabus', event)" onmouseover="showBaloon(2, 'Carabus', event)" target="_blank">
Carabus
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Eucarabus" onmouseout="hideBaloonEvent('Eucarabus', event)" onmouseover="showBaloon(2, 'Eucarabus', event)" target="_blank">
(Eucarabus)
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Carabus%20%28Eucarabus%29%20deyrollei" onmouseout="hideBaloonEvent('Carabus_(Eucarabus)_deyrollei', event)" onmouseover="showBaloon(2, 'Carabus_(Eucarabus)_deyrollei', event)" target="_blank">
deyrollei
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Lugo, Spain
</td>
<td class="cell_3" colspan="1">
1553BG
</td>
<td class="cell_4" colspan="1">
JF778782
</td>
<td class="cell_5" colspan="1">
JF778799
</td>
<td class="cell_6" colspan="1">
JF778815
</td>
<td class="cell_7" colspan="1">
JF778832
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Carabus" onmouseout="hideBaloonEvent('Carabus', event)" onmouseover="showBaloon(2, 'Carabus', event)" target="_blank">
Carabus
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Limnocarabaus" onmouseout="hideBaloonEvent('Limnocarabaus', event)" onmouseover="showBaloon(2, 'Limnocarabaus', event)" target="_blank">
(Limnocarabaus)
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Carabus%20%28Limnocarabaus%29%20clathratus" onmouseout="hideBaloonEvent('Carabus_(Limnocarabaus)_clathratus', event)" onmouseover="showBaloon(2, 'Carabus_(Limnocarabaus)_clathratus', event)" target="_blank">
clathratus
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Susuz, Turkey
</td>
<td class="cell_3" colspan="1">
1600BG
</td>
<td class="cell_4" colspan="1">
JF778783
</td>
<td class="cell_5" colspan="1">
JF778800
</td>
<td class="cell_6" colspan="1">
JF778816
</td>
<td class="cell_7" colspan="1">
JF778833
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Dixus" onmouseout="hideBaloonEvent('Dixus', event)" onmouseover="showBaloon(2, 'Dixus', event)" target="_blank">
Dixus
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Dixus%20capito" onmouseout="hideBaloonEvent('Dixus_capito', event)" onmouseover="showBaloon(2, 'Dixus_capito', event)" target="_blank">
capito
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Albacete, Spain
</td>
<td class="cell_3" colspan="1">
1578BG
</td>
<td class="cell_4" colspan="1">
JF778784
</td>
<td class="cell_5" colspan="1">
N/A
</td>
<td class="cell_6" colspan="1">
JF778817
</td>
<td class="cell_7" colspan="1">
JF778834
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Pseudotrechus" onmouseout="hideBaloonEvent('Pseudotrechus', event)" onmouseover="showBaloon(2, 'Pseudotrechus', event)" target="_blank">
Pseudotrechus
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Pseudotrechus%20mutilatus" onmouseout="hideBaloonEvent('Pseudotrechus_mutilatus', event)" onmouseover="showBaloon(2, 'Pseudotrechus_mutilatus', event)" target="_blank">
mutilatus
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Cádiz, Spain
</td>
<td class="cell_3" colspan="1">
36_EN
</td>
<td class="cell_4" colspan="1">
JF778785
</td>
<td class="cell_5" colspan="1">
JF778801
</td>
<td class="cell_6" colspan="1">
JF778818
</td>
<td class="cell_7" colspan="1">
JF778835
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Licinus" onmouseout="hideBaloonEvent('Licinus', event)" onmouseover="showBaloon(2, 'Licinus', event)" target="_blank">
Licinus
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Licinus%20punctatulus" onmouseout="hideBaloonEvent('Licinus_punctatulus', event)" onmouseover="showBaloon(2, 'Licinus_punctatulus', event)" target="_blank">
punctatulus
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Alicante, Spain
</td>
<td class="cell_3" colspan="1">
1582BG
</td>
<td class="cell_4" colspan="1">
JF778786
</td>
<td class="cell_5" colspan="1">
JF778802
</td>
<td class="cell_6" colspan="1">
JF778819
</td>
<td class="cell_7" colspan="1">
JF778836
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Elaphropus" onmouseout="hideBaloonEvent('Elaphropus', event)" onmouseover="showBaloon(2, 'Elaphropus', event)" target="_blank">
Elaphropus
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Tachyura" onmouseout="hideBaloonEvent('Tachyura', event)" onmouseover="showBaloon(2, 'Tachyura', event)" target="_blank">
(Tachyura)
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Elaphropus%20%28Tachyura%29%20parvulus" onmouseout="hideBaloonEvent('Elaphropus_(Tachyura)_parvulus', event)" onmouseover="showBaloon(2, 'Elaphropus_(Tachyura)_parvulus', event)" target="_blank">
parvulus
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Pays Zaer Zaine, Morocco
</td>
<td class="cell_3" colspan="1">
64_EN
</td>
<td class="cell_4" colspan="1">
JF778787
</td>
<td class="cell_5" colspan="1">
JF778803
</td>
<td class="cell_6" colspan="1">
JF778820
</td>
<td class="cell_7" colspan="1">
JF778837
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Bembidion" onmouseout="hideBaloonEvent('Bembidion', event)" onmouseover="showBaloon(2, 'Bembidion', event)" target="_blank">
Bembidion
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Peryphus" onmouseout="hideBaloonEvent('Peryphus', event)" onmouseover="showBaloon(2, 'Peryphus', event)" target="_blank">
(Peryphus)
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Bembidion%20%28Peryphus%29%20hispanicum" onmouseout="hideBaloonEvent('Bembidion_(Peryphus)_hispanicum', event)" onmouseover="showBaloon(2, 'Bembidion_(Peryphus)_hispanicum', event)" target="_blank">
hispanicum
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Pays Zaer Zaine, Morocco
</td>
<td class="cell_3" colspan="1">
62_EN
</td>
<td class="cell_4" colspan="1">
N/A
</td>
<td class="cell_5" colspan="1">
JF778804
</td>
<td class="cell_6" colspan="1">
JF778821
</td>
<td class="cell_7" colspan="1">
JF778838
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Bembidion" onmouseout="hideBaloonEvent('Bembidion', event)" onmouseover="showBaloon(2, 'Bembidion', event)" target="_blank">
Bembidion
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Emphanes" onmouseout="hideBaloonEvent('Emphanes', event)" onmouseover="showBaloon(2, 'Emphanes', event)" target="_blank">
(Emphanes)
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Bembidion%20%28Emphanes%29%20latiplaga" onmouseout="hideBaloonEvent('Bembidion_(Emphanes)_latiplaga', event)" onmouseover="showBaloon(2, 'Bembidion_(Emphanes)_latiplaga', event)" target="_blank">
latiplaga
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Pays Zaer Zaine, Morocco
</td>
<td class="cell_3" colspan="1">
65_EN
</td>
<td class="cell_4" colspan="1">
JF778788
</td>
<td class="cell_5" colspan="1">
N/A
</td>
<td class="cell_6" colspan="1">
JF778822
</td>
<td class="cell_7" colspan="1">
JF778839
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Perileptus" onmouseout="hideBaloonEvent('Perileptus', event)" onmouseover="showBaloon(2, 'Perileptus', event)" target="_blank">
Perileptus
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Perileptus%20aerolatus" onmouseout="hideBaloonEvent('Perileptus_aerolatus', event)" onmouseover="showBaloon(2, 'Perileptus_aerolatus', event)" target="_blank">
aerolatus
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Agadir, Morocco
</td>
<td class="cell_3" colspan="1">
MNHN-AF113
</td>
<td class="cell_4" colspan="1">
GQ293688
</td>
<td class="cell_5" colspan="1">
FR729593
</td>
<td class="cell_6" colspan="1">
GQ293625
</td>
<td class="cell_7" colspan="1">
GQ293503
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Trechus" onmouseout="hideBaloonEvent('Trechus', event)" onmouseover="showBaloon(2, 'Trechus', event)" target="_blank">
Trechus
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Trechus%20quadristriatus" onmouseout="hideBaloonEvent('Trechus_quadristriatus', event)" onmouseover="showBaloon(2, 'Trechus_quadristriatus', event)" target="_blank">
quadristriatus
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Huesca, Spain
</td>
<td class="cell_3" colspan="1">
MNHN-AF96
</td>
<td class="cell_4" colspan="1">
FR733908
</td>
<td class="cell_5" colspan="1">
GQ293743
</td>
<td class="cell_6" colspan="1">
GQ293619
</td>
<td class="cell_7" colspan="1">
GQ293534
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Typhloreicheia" onmouseout="hideBaloonEvent('Typhloreicheia', event)" onmouseover="showBaloon(2, 'Typhloreicheia', event)" target="_blank">
Typhloreicheia
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Typhloreicheia%20laurentii" onmouseout="hideBaloonEvent('Typhloreicheia_laurentii', event)" onmouseover="showBaloon(2, 'Typhloreicheia_laurentii', event)" target="_blank">
laurentii
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Sardinia, Italy
</td>
<td class="cell_3" colspan="1">
56_EN
</td>
<td class="cell_4" colspan="1">
JF778789
</td>
<td class="cell_5" colspan="1">
JF778805
</td>
<td class="cell_6" colspan="1">
JF778823
</td>
<td class="cell_7" colspan="1">
JF778840
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Dyschiriodes" onmouseout="hideBaloonEvent('Dyschiriodes', event)" onmouseover="showBaloon(2, 'Dyschiriodes', event)" target="_blank">
Dyschiriodes
</a>
</span>
</span>
</i>
sp.
</td>
<td class="cell_2" colspan="1">
Pays Zaer Zaine, Morocco
</td>
<td class="cell_3" colspan="1">
63_EN
</td>
<td class="cell_4" colspan="1">
JF778790
</td>
<td class="cell_5" colspan="1">
JF778806
</td>
<td class="cell_6" colspan="1">
JF778824
</td>
<td class="cell_7" colspan="1">
JF778841
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Nebria" onmouseout="hideBaloonEvent('Nebria', event)" onmouseover="showBaloon(2, 'Nebria', event)" target="_blank">
Nebria
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Nebria%20salina" onmouseout="hideBaloonEvent('Nebria_salina', event)" onmouseover="showBaloon(2, 'Nebria_salina', event)" target="_blank">
salina
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Albacete, Spain
</td>
<td class="cell_3" colspan="1">
1579BG
</td>
<td class="cell_4" colspan="1">
JF778791
</td>
<td class="cell_5" colspan="1">
JF778807
</td>
<td class="cell_6" colspan="1">
JF778825
</td>
<td class="cell_7" colspan="1">
JF778842
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Ildobates" onmouseout="hideBaloonEvent('Ildobates', event)" onmouseover="showBaloon(2, 'Ildobates', event)" target="_blank">
Ildobates
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Ildobates%20neboti" onmouseout="hideBaloonEvent('Ildobates_neboti', event)" onmouseover="showBaloon(2, 'Ildobates_neboti', event)" target="_blank">
neboti
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Castellón, Spain
</td>
<td class="cell_3" colspan="1">
MNCN-6409
</td>
<td class="cell_4" colspan="1">
JF778792
</td>
<td class="cell_5" colspan="1">
JF778808
</td>
<td class="cell_6" colspan="1">
AM051084
</td>
<td class="cell_7" colspan="1">
DQ130051
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Drypta" onmouseout="hideBaloonEvent('Drypta', event)" onmouseover="showBaloon(2, 'Drypta', event)" target="_blank">
Drypta
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Drypta%20dentata" onmouseout="hideBaloonEvent('Drypta_dentata', event)" onmouseover="showBaloon(2, 'Drypta_dentata', event)" target="_blank">
dentata
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Ciudad Real, Spain
</td>
<td class="cell_3" colspan="1">
98_EN
</td>
<td class="cell_4" colspan="1">
N/A
</td>
<td class="cell_5" colspan="1">
JF778809
</td>
<td class="cell_6" colspan="1">
N/A
</td>
<td class="cell_7" colspan="1">
N/A
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Zuphium" onmouseout="hideBaloonEvent('Zuphium', event)" onmouseover="showBaloon(2, 'Zuphium', event)" target="_blank">
Zuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Zuphium%20olens" onmouseout="hideBaloonEvent('Zuphium_olens', event)" onmouseover="showBaloon(2, 'Zuphium_olens', event)" target="_blank">
olens
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Murcia, Spain
</td>
<td class="cell_3" colspan="1">
97_EN
</td>
<td class="cell_4" colspan="1">
JF778793
</td>
<td class="cell_5" colspan="1">
N/A
</td>
<td class="cell_6" colspan="1">
JF778826
</td>
<td class="cell_7" colspan="1">
JF778843
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20cf.%20baeticum" onmouseout="hideBaloonEvent('Parazuphium_cf__baeticum', event)" onmouseover="showBaloon(2, 'Parazuphium_cf__baeticum', event)" target="_blank">
cf.
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20cf.%20baeticum" onmouseout="hideBaloonEvent('Parazuphium_cf__baeticum', event)" onmouseover="showBaloon(2, 'Parazuphium_cf__baeticum', event)" target="_blank">
baeticum
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Castellón, Spain
</td>
<td class="cell_3" colspan="1">
87_EN
</td>
<td class="cell_4" colspan="1">
JF778794
</td>
<td class="cell_5" colspan="1">
JF778810
</td>
<td class="cell_6" colspan="1">
JF778827
</td>
<td class="cell_7" colspan="1">
JF778844
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" onmouseout="hideBaloonEvent('Parazuphium_aguilerai', event)" onmouseover="showBaloon(2, 'Parazuphium_aguilerai', event)" target="_blank">
aguilerai
</a>
</span>
</span>
</i>
</td>
<td class="cell_2" colspan="1">
Tanger, Morocco
</td>
<td class="cell_3" colspan="1">
31_EN
</td>
<td class="cell_4" colspan="1">
JF778795
</td>
<td class="cell_5" colspan="1">
JF778811
</td>
<td class="cell_6" colspan="1">
JF778828
</td>
<td class="cell_7" colspan="1">
JF778845
</td>
</tr>
</tbody>
</table>
</div>
<a name="phylogeneticmethods">
</a>
<span class="node_title1">
Phylogenetic methods
</span>
<p>
Bayesian phylogenetic analyses (BA) were performed with MrBayes v.3.1. (
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B4')" onmouseover="showBaloon(1, 'B4', event)">
Huelsenbeck and Ronquist 2001
</span>
,
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B18')" onmouseover="showBaloon(1, 'B18', event)">
Ronquist and Huelsenbeck 2003
</span>
),
partitioning by gene with a GTR+G model applied to each partition. Two
independent runs of 20, 000, 000 generations were conducted, each with
three hot and one cold chain, whereby trees were sampled every 100
generations. Sampled trees were analysed with Tracer v.1.5 (
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B16')" onmouseover="showBaloon(1, 'B16', event)">
Rambaut and Drummond 2007
</span>
)
and their half compact consensus tree was calculated with a burning
value of 10% with node posterior probabilities used as support values,
checking for an appropriate degree of convergence between chains with
the effective sample size in Tracer v.1.5. MrBayes was run on-line at
the freely available computational service of Bioportal
(www.bioportal.uio.no). Trees were visualized in FigTree v.1.3.1 (
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B15')" onmouseover="showBaloon(1, 'B15', event)">
Rambaut 2008
</span>
).
</p>
<a name="speciestreatment">
</a>
<span class="node_title">
Species treatment
</span>
<div class="taxonTreatmentsHolder secHolder">
<div class="taxonTreatmentRow node_taxon-treatment">
<div class="taxonNomenclature">
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
<i>
Parazuphium
</i>
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20Parazuphium%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20aguilerai%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20" onmouseout="hideBaloonEvent('Parazuphium_aguilerai', event)" onmouseover="showBaloon(2, 'Parazuphium_aguilerai', event)" target="_blank">
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20Parazuphium%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20aguilerai%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20" onmouseout="hideBaloonEvent('Parazuphium_aguilerai', event)" onmouseover="showBaloon(2, 'Parazuphium_aguilerai', event)" target="_blank">
</a>
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</a>
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<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20Parazuphium%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20aguilerai%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20" onmouseout="hideBaloonEvent('Parazuphium_aguilerai', event)" onmouseover="showBaloon(2, 'Parazuphium_aguilerai', event)" target="_blank">
</a>
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</a>
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</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20Parazuphium%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20aguilerai%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20%20" onmouseout="hideBaloonEvent('Parazuphium_aguilerai', event)" onmouseover="showBaloon(2, 'Parazuphium_aguilerai', event)" target="_blank">
<i>
aguilerai
</i>
</a>
</span>
</span>
Andújar, Hernando & Ribera
sp. n.
<p class="node_object_id">
<a href="http://zoobank.org/?lsid=urn:lsid:zoobank.org:act:B4718866-DA9E-4096-9291-A38E90FD7A0A" target="_blank">
urn:lsid:zoobank.org:act:B4718866-DA9E-4096-9291-A38E90FD7A0A
</a>
</p>
<p class="node_object_id">
<a href="http://species-id.net/wiki/Parazuphium_aguilerai" target="_blank">
http://species-id.net/wiki/Parazuphium_aguilerai
</a>
</p>
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F1')" onmouseover="showBaloon(3, 'F1', event)">
Figs 1
</span>
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F2')" onmouseover="showBaloon(3, 'F2', event)">
</span>
–
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F3')" onmouseover="showBaloon(3, 'F3', event)">
3
</span>
</div>
<div class="taxonTreatmentSec inlineSec">
<span class="node_title">
Type locality.
</span>
<p>
Souk-Khemis-des-Anjra, Tetuan, Morocco (
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F4')" onmouseover="showBaloon(3, 'F4', event)">
Fig. 4
</span>
).
</p>
<div class="unfloat">
</div>
</div>
<div class="taxonTreatmentSec inlineSec">
<span class="node_title">
Type specimen.
</span>
<p>
Holotype: 1♂, “MOROCCO 28-III-2008 /
Souk-Khemis-des-Anjra, Tetuan / 123m N35°43'18" W5°31'23" / Andújar,
Hernando, Ribera & Aguilera leg."; voucher number label “31_EN";
plus red holotype label. Type specimen mounted in DMHF in a transparent
acetate label, genitalia dissected and mounted in DMHF in a separate
label pinned with the specimen. Deposited in the Museu de Ciències
Naturals de Barcelona (MCNB), DNA aliquots deposited in the IBE (CSIC)
and Univ. Murcia (ZAFUMU col.).
</p>
<div class="unfloat">
</div>
</div>
<div class="taxonTreatmentSec inlineSec">
<span class="node_title">
Diagnosis.
</span>
<p>
Total Length 2.7 mm (from apex of mandible to apex of elytra). Body depressed, flattened, light brown (
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F1')" onmouseover="showBaloon(3, 'F1', event)">
Fig. 1
</span>
). Eyes absent (
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F2')" onmouseover="showBaloon(3, 'F2', event)">
Fig. 2b
</span>
). First antennomere (0.41mm) as long as antennomeres 2–4 combined (0.37 mm) (
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F2')" onmouseover="showBaloon(3, 'F2', event)">
Fig. 2e
</span>
). Pronotum cordiform (
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F1')" onmouseover="showBaloon(3, 'F1', event)">
Fig. 1
</span>
). Elytra flat, not fully covering abdomen (
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F1')" onmouseover="showBaloon(3, 'F1', event)">
Figs 1
</span>
,
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F2')" onmouseover="showBaloon(3, 'F2', event)">
2a
</span>
). Umbilicate lateral series of elytra with 5+5 spatuliform setae (
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F2')" onmouseover="showBaloon(3, 'F2', event)">
Figs 2f-h
</span>
). Apex of elytra divergent (
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F1')" onmouseover="showBaloon(3, 'F1', event)">
Figs 1
</span>
,
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F2')" onmouseover="showBaloon(3, 'F2', event)">
2a
</span>
). Metafemora with an acute tooth on interior margin (
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F2')" onmouseover="showBaloon(3, 'F2', event)">
Fig. 2m
</span>
).
</p>
<div class="imgFloatLeft">
<div class="graphicHolder">
<div class="graphicImg">
<a href="export.php_files/ZooKeys-103-049-g001.jpg" target="_blank">
<img alt="" src="export.php_files/ZooKeys-103-049-g001.jpg"/>
</a>
</div>
<div class="graphicCaptions">
<span class="graphicCaption">
</span>
<span class="graphicDescription">
</span>
</div>
<div class="unfloat">
</div>
</div>
</div>
<div class="ImgTextFloatRight">
<span class="node_label">
Figure 1.
</span>
<p>
Line drawing ofhabitus of
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" onmouseout="hideBaloonEvent('Parazuphium_aguilerai', event)" onmouseover="showBaloon(2, 'Parazuphium_aguilerai', event)" target="_blank">
aguilerai
</a>
</span>
</span>
</i>
sp. n. Total length 2.7 mm.
</p>
</div>
<div class="unfloat">
</div>
<div class="figureHolder" id="fig_F1">
<div class="imgFloatLeft">
<div class="graphicHolder">
<div class="graphicImg">
<a href="export.php_files/ZooKeys-103-049-g001.jpg" target="_blank">
<img alt="" src="export.php_files/ZooKeys-103-049-g001.jpg"/>
</a>
</div>
<div class="graphicCaptions">
<span class="graphicCaption">
</span>
<span class="graphicDescription">
</span>
</div>
<div class="unfloat">
</div>
</div>
</div>
<div class="ImgTextFloatRight">
<span class="node_label">
Figure 1.
</span>
<p>
Line drawing ofhabitus of
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" target="_blank">
aguilerai
</a>
</span>
</span>
</i>
sp. n. Total length 2.7 mm.
</p>
</div>
<div class="unfloat">
</div>
</div>
<div class="unfloat">
</div>
</div>
<div class="taxonTreatmentSec inlineSec">
<span class="node_title">
Description.
</span>
<p>
Length of holotype: 2.7 mm. Body depressed,
flattened and depigmented, light brown. Surface microreticulate, with
mesh pattern regular polygonal (observed on the dried specimen) and
scattered short setae.
</p>
<p>
<i>
Head
</i>
(
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F1')" onmouseover="showBaloon(3, 'F1', event)">
Fig. 1
</span>
) with trapezoidal shape. No trace of eyes or ocular scars (
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F2')" onmouseover="showBaloon(3, 'F2', event)">
Fig. 2b
</span>
).
Length of head (from apex of mandible to base) 0.63 mm; maximum width
close to base (0.51 mm). Surface microreticulate, microlines deeper on
sides. Neck pedunculate. With three long setae, two lateral and one
basal. Appendages: antennae (
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F2')" onmouseover="showBaloon(3, 'F2', event)">
Fig. 2e
</span>
)
with first antennomere (0.41mm) as long as total length of antennomeres
2–3-4 together (0.37 mm); second antennomere pedunculate (0.1 mm),
slightly shorter than third (0.13 mm) and fourth (0.14 mm); from fifth
to tenth with same length (0.16–0.17 mm); last antennomere longer
(0.23mm). Antennomeres from 3° to 11° cylindrical. Labial and maxillary
palpi as in
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F2')" onmouseover="showBaloon(3, 'F2', event)">
Figs 2c, d
</span>
.
</p>
<div class="imgFloatLeft">
<div class="graphicHolder">
<div class="graphicImg">
<a href="export.php_files/ZooKeys-103-049-g002.jpg" target="_blank">
<img alt="" src="export.php_files/ZooKeys-103-049-g002.jpg"/>
</a>
</div>
<div class="graphicCaptions">
<span class="graphicCaption">
</span>
<span class="graphicDescription">
</span>
</div>
<div class="unfloat">
</div>
</div>
</div>
<div class="ImgTextFloatRight">
<span class="node_label">
Figure 2.
</span>
<p>
Photographic images of
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" onmouseout="hideBaloonEvent('Parazuphium_aguilerai', event)" onmouseover="showBaloon(2, 'Parazuphium_aguilerai', event)" target="_blank">
aguilerai
</a>
</span>
</span>
</i>
sp. n.
<b>
A
</b>
whole specimen
<b>
B
</b>
head in dorso-lateral view
<b>
C
</b>
labial palpus; (d), maxillary palpus
<b>
C
</b>
antenna
<b>
F
</b>
margin of left elytron in lateral view
<b>
G
</b>
margin of right elytron, detail for anterior umbilicate setae, numbers 1 to 5
<b>
H
</b>
margin of right elytron, detail of posterior umbilicate setae, numbers 6 to10, arrows over them point other smaller setae
<b>
I–N
</b>
details of anterior, median and posterior legs respectively.
</p>
</div>
<div class="unfloat">
</div>
<div class="figureHolder" id="fig_F2">
<div class="imgFloatLeft">
<div class="graphicHolder">
<div class="graphicImg">
<a href="export.php_files/ZooKeys-103-049-g002.jpg" target="_blank">
<img alt="" src="export.php_files/ZooKeys-103-049-g002.jpg"/>
</a>
</div>
<div class="graphicCaptions">
<span class="graphicCaption">
</span>
<span class="graphicDescription">
</span>
</div>
<div class="unfloat">
</div>
</div>
</div>
<div class="ImgTextFloatRight">
<span class="node_label">
Figure 2.
</span>
<p>
Photographic images of
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" target="_blank">
aguilerai
</a>
</span>
</span>
</i>
sp. n.
<b>
A
</b>
whole specimen
<b>
B
</b>
head in dorso-lateral view
<b>
C
</b>
labial palpus; (d), maxillary palpus
<b>
C
</b>
antenna
<b>
F
</b>
margin of left elytron in lateral view
<b>
G
</b>
margin of right elytron, detail for anterior umbilicate setae, numbers 1 to 5
<b>
H
</b>
margin of right elytron, detail of posterior umbilicate setae, numbers 6 to10, arrows over them point other smaller setae
<b>
I–N
</b>
details of anterior, median and posterior legs respectively.
</p>
</div>
<div class="unfloat">
</div>
</div>
<p>
<i>
Pronotum
</i>
cordiform (
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F1')" onmouseover="showBaloon(3, 'F1', event)">
Fig. 1
</span>
),
longer (0.60 mm) than wide (0.51–0.27 mm), maximum width (0.51 mm)
close to anterior angles, almost double minimum width (0.27 mm), at
the posterior angles. Anterior angles obtuse, rounded. Anterior margin
regularly convex. Median line apparent, marked with two depressions.
Two lateral setae at anterior and posterior angles. Lateral margin
sinuate before posterior angles.
</p>
<p>
<i>
Elytra
</i>
(
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F1')" onmouseover="showBaloon(3, 'F1', event)">
Figs 1
</span>
,
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F2')" onmouseover="showBaloon(3, 'F2', event)">
2a
</span>
)
flattened, short, not totally covering abdomen, wider apically
(maximum width, 0.90mm, close to apex); width at humeral angle 0.65mm.
Punctuation forming longitudinal series, more evident at basal third,
disappearing towards apex. Entire surface with short pubescence.
Anterior umbilicate series with 5 spatuliform setae (
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F2')" onmouseover="showBaloon(3, 'F2', event)">
Figs 2f-g
</span>
, numbers 1–5), deeply inserted in small marginal indentations, with some other minor setae over them
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F2')" onmouseover="showBaloon(3, 'F2', event)">
(Fig. 2g
</span>
, arrows). Posterior umbilicate series with 5 spatuliform setae, the last one just before apex (
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F2')" onmouseover="showBaloon(3, 'F2', event)">
Fig. 2h
</span>
, numbers 6–10), with three smaller setae over them (
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F2')" onmouseover="showBaloon(3, 'F2', event)">
Fig. 2h
</span>
, arrows). Margin of elytra from 5° umbilical anterior to 2° umbilical posterior seta with a marginal carina (
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F2')" onmouseover="showBaloon(3, 'F2', event)">
Fig. 2f
</span>
). Apices divergent (
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F1')" onmouseover="showBaloon(3, 'F1', event)">
Figs 1
</span>
,
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F2')" onmouseover="showBaloon(3, 'F2', event)">
2a
</span>
).
</p>
<p>
<i>
Legs
</i>
. Pro- and meso-femora dilated proximally, forming an obtuse interior angle (
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F2')" onmouseover="showBaloon(3, 'F2', event)">
Figs 2i, k
</span>
). Metafemora with a strong acute tooth on the interior margin (
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F2')" onmouseover="showBaloon(3, 'F2', event)">
Fig. 2m
</span>
). Front tibia with antennal cleaner (toilette organ), as reported in other species of the genus (
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F2')" onmouseover="showBaloon(3, 'F2', event)">
Fig. 2i
</span>
).
Meta-tibia long and straight, with an internal spine at apex. Meso and
meta tibiae with a circle of seta round the apex. Pro-tarsomeres 1–4
dilated (
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F2')" onmouseover="showBaloon(3, 'F2', event)">
Fig. 2j
</span>
). First meso- and meta-tarsomeres as long as 2° to 4° combined (
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F2')" onmouseover="showBaloon(3, 'F2', event)">
Figs 2l, n
</span>
). Fourth tarsomere cordiform. Trochanters without tooth or any other special structure.
</p>
<p>
<i>
Aedeagus
</i>
. Median lobe as in
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F3')" onmouseover="showBaloon(3, 'F3', event)">
Fig. 3
</span>
,
short and robust with a ventral constriction between the basal and the
distal part as described for the genus. Basal margin arcuate,
bisinuate, with the apex rounded. Internal sac with two small
sclerites. Parameres asymmetric, as in other species of the genus.
</p>
<div class="imgFloatLeft">
<div class="graphicHolder">
<div class="graphicImg">
<a href="export.php_files/ZooKeys-103-049-g003.jpg" target="_blank">
<img alt="" src="export.php_files/ZooKeys-103-049-g003.jpg"/>
</a>
</div>
<div class="graphicCaptions">
<span class="graphicCaption">
</span>
<span class="graphicDescription">
</span>
</div>
<div class="unfloat">
</div>
</div>
</div>
<div class="ImgTextFloatRight">
<span class="node_label">
Figure 3.
</span>
<p>
Photographic image of median lobe of
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" onmouseout="hideBaloonEvent('Parazuphium_aguilerai', event)" onmouseover="showBaloon(2, 'Parazuphium_aguilerai', event)" target="_blank">
aguilerai
</a>
</span>
</span>
</i>
sp. n. Left lateral aspect.
</p>
</div>
<div class="unfloat">
</div>
<div class="figureHolder" id="fig_F3">
<div class="imgFloatLeft">
<div class="graphicHolder">
<div class="graphicImg">
<a href="export.php_files/ZooKeys-103-049-g003.jpg" target="_blank">
<img alt="" src="export.php_files/ZooKeys-103-049-g003.jpg"/>
</a>
</div>
<div class="graphicCaptions">
<span class="graphicCaption">
</span>
<span class="graphicDescription">
</span>
</div>
<div class="unfloat">
</div>
</div>
</div>
<div class="ImgTextFloatRight">
<span class="node_label">
Figure 3.
</span>
<p>
Photographic image of median lobe of
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" target="_blank">
aguilerai
</a>
</span>
</span>
</i>
sp. n. Left lateral aspect.
</p>
</div>
<div class="unfloat">
</div>
</div>
<div class="unfloat">
</div>
</div>
<div class="taxonTreatmentSec inlineSec">
<span class="node_title">
Habitat.
</span>
<p>
The single known specimen of
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" onmouseout="hideBaloonEvent('Parazuphium_aguilerai', event)" onmouseover="showBaloon(2, 'Parazuphium_aguilerai', event)" target="_blank">
aguilerai
</a>
</span>
</span>
</i>
sp. n. was found under a large, deeply buried boulder, in the humid soil on a hillside with herbaceous vegetation (
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Chamaerops" onmouseout="hideBaloonEvent('Chamaerops', event)" onmouseover="showBaloon(2, 'Chamaerops', event)" target="_blank">
Chamaerops
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Chamaerops%20humilis" onmouseout="hideBaloonEvent('Chamaerops_humilis', event)" onmouseover="showBaloon(2, 'Chamaerops_humilis', event)" target="_blank">
humilis
</a>
</span>
</span>
</i>
,
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Nerium" onmouseout="hideBaloonEvent('Nerium', event)" onmouseover="showBaloon(2, 'Nerium', event)" target="_blank">
Nerium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Nerium%20oleander" onmouseout="hideBaloonEvent('Nerium_oleander', event)" onmouseover="showBaloon(2, 'Nerium_oleander', event)" target="_blank">
oleander
</a>
</span>
</span>
</i>
and
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Pistacia" onmouseout="hideBaloonEvent('Pistacia', event)" onmouseover="showBaloon(2, 'Pistacia', event)" target="_blank">
Pistacia
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Pistacia%20lentiscus" onmouseout="hideBaloonEvent('Pistacia_lentiscus', event)" onmouseover="showBaloon(2, 'Pistacia_lentiscus', event)" target="_blank">
lentiscus
</a>
</span>
</span>
</i>
,
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F4')" onmouseover="showBaloon(3, 'F4', event)">
Fig. 4
</span>
). The same sample included some endogean ants (
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Leptanilla" onmouseout="hideBaloonEvent('Leptanilla', event)" onmouseover="showBaloon(2, 'Leptanilla', event)" target="_blank">
Leptanilla
</a>
</span>
</span>
</i>
sp,
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Amblyopone" onmouseout="hideBaloonEvent('Amblyopone', event)" onmouseover="showBaloon(2, 'Amblyopone', event)" target="_blank">
Amblyopone
</a>
</span>
</span>
</i>
sp.) and remains of anendogean weevil,
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Torneuma" onmouseout="hideBaloonEvent('Torneuma', event)" onmouseover="showBaloon(2, 'Torneuma', event)" target="_blank">
Torneuma
</a>
</span>
</span>
</i>
sp. (
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Curculionidae" onmouseout="hideBaloonEvent('Curculionidae', event)" onmouseover="showBaloon(2, 'Curculionidae', event)" target="_blank">
Curculionidae
</a>
</span>
</span>
,
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Cryptorhynchinae" onmouseout="hideBaloonEvent('Cryptorhynchinae', event)" onmouseover="showBaloon(2, 'Cryptorhynchinae', event)" target="_blank">
Cryptorhynchinae
</a>
</span>
</span>
).
</p>
<div class="imgFloatLeft">
<div class="graphicHolder">
<div class="graphicImg">
<a href="export.php_files/ZooKeys-103-049-g004.jpg" target="_blank">
<img alt="" src="export.php_files/ZooKeys-103-049-g004.jpg"/>
</a>
</div>
<div class="graphicCaptions">
<span class="graphicCaption">
</span>
<span class="graphicDescription">
</span>
</div>
<div class="unfloat">
</div>
</div>
</div>
<div class="ImgTextFloatRight">
<span class="node_label">
Figure 4.
</span>
<p>
Habitat of
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" onmouseout="hideBaloonEvent('Parazuphium_aguilerai', event)" onmouseover="showBaloon(2, 'Parazuphium_aguilerai', event)" target="_blank">
aguilerai
</a>
</span>
</span>
</i>
sp. n.
</p>
</div>
<div class="unfloat">
</div>
<div class="figureHolder" id="fig_F4">
<div class="imgFloatLeft">
<div class="graphicHolder">
<div class="graphicImg">
<a href="export.php_files/ZooKeys-103-049-g004.jpg" target="_blank">
<img alt="" src="export.php_files/ZooKeys-103-049-g004.jpg"/>
</a>
</div>
<div class="graphicCaptions">
<span class="graphicCaption">
</span>
<span class="graphicDescription">
</span>
</div>
<div class="unfloat">
</div>
</div>
</div>
<div class="ImgTextFloatRight">
<span class="node_label">
Figure 4.
</span>
<p>
Habitat of
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" target="_blank">
aguilerai
</a>
</span>
</span>
</i>
sp. n.
</p>
</div>
<div class="unfloat">
</div>
</div>
<div class="unfloat">
</div>
</div>
<div class="taxonTreatmentSec inlineSec">
<span class="node_title">
Etymology.
</span>
<p>
The specific epithet is a Latinized eponym,
genitive case, based on the name of our late friend Pedro Aguilera,
who collected the specimen with us during his last trip to Morocco.
</p>
<div class="unfloat">
</div>
</div>
<div class="taxonTreatmentSec inlineSec">
<span class="node_title">
Recognition and comparisons.
</span>
<p>
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" onmouseout="hideBaloonEvent('Parazuphium_aguilerai', event)" onmouseover="showBaloon(2, 'Parazuphium_aguilerai', event)" target="_blank">
aguilerai
</a>
</span>
</span>
</i>
sp. n. can be clearly distinguished from any other species of the genus
through the combinations of the following characters: lack of eyes,
reduced size (2.7 mm), length and proportions of 2°, 3° and 4°
antennomeres (0.1, 0.13 and 0.14mm respectively) and the presence of a
tooth on metafemora.
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20feloi" onmouseout="hideBaloonEvent('Parazuphium_feloi', event)" onmouseover="showBaloon(2, 'Parazuphium_feloi', event)" target="_blank">
feloi
</a>
</span>
</span>
</i>
from the Canary Islands is also anophthalmous, but it is larger than
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" onmouseout="hideBaloonEvent('Parazuphium_aguilerai', event)" onmouseover="showBaloon(2, 'Parazuphium_aguilerai', event)" target="_blank">
aguilerai
</a>
</span>
</span>
</i>
sp. n. and without a tooth on the hind femora (
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B11')" onmouseover="showBaloon(1, 'B11', event)">
Machado 1998
</span>
).
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20ramirezi" onmouseout="hideBaloonEvent('Parazuphium_ramirezi', event)" onmouseover="showBaloon(2, 'Parazuphium_ramirezi', event)" target="_blank">
ramirezi
</a>
</span>
</span>
</i>
J. and E. Vives from south Spain shows the same tooth on the metafemora, but is also larger, and with reduced eyes (
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B22')" onmouseover="showBaloon(1, 'B22', event)">
Vives and Vives 1976
</span>
). There are also some differences in the shape of the head and pronotum: in
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" onmouseout="hideBaloonEvent('Parazuphium_aguilerai', event)" onmouseover="showBaloon(2, 'Parazuphium_aguilerai', event)" target="_blank">
aguilerai
</a>
</span>
</span>
</i>
sp. n. the head is more parallel-sided, the anterior angles of the
pronotum are less rounded, and the anterior margin not straight.
</p>
<div class="unfloat">
</div>
</div>
</div>
</div>
<a name="identificationkey">
</a>
<span class="node_title">
Identification key
</span>
<p>
Key to adults of the West Mediterranean and Macaronesian
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
</span>
</span>
</i>
species, modified from
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B1')" onmouseover="showBaloon(1, 'B1', event)">
Antoine (1962)
</span>
and
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B5')" onmouseover="showBaloon(1, 'B5', event)">
Hürka (1982)
</span>
:
</p>
<div class="tableWrapHolder keyTable">
<div class="tableDiv" id="table_">
<table class="innerTable Table">
<tbody>
<tr>
<td class="cell_1" colspan="1">
1
</td>
<td class="cell_2" colspan="1">
Eyeless
</td>
<td class="cell_3" colspan="1">
2
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
–
</td>
<td class="cell_2" colspan="1">
With eyes
</td>
<td class="cell_3" colspan="1">
3
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
2
</td>
<td class="cell_2" colspan="1">
Third antennal segment only slightly
longer than 2nd and slightly shorter than 4th, anterior margin of
pronotum trapezoidal, presence of a tooth on metafemora. Length 2.7mm.
North Morocco
</td>
<td class="cell_3" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" target="_blank">
aguilerai
</a>
</span>
</span>
</i>
sp. n.
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
–
</td>
<td class="cell_2" colspan="1">
Third antennal segment more than twice
longer than 2nd and similar to 4th. Anterior margin of pronotum
bisinuate, without tooth on metafemora. Length 4.9–5.1mm. Canary
Islands
</td>
<td class="cell_3" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20feloi" target="_blank">
feloi
</a>
</span>
</span>
</i>
Machado
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
3
</td>
<td class="cell_2" colspan="1">
Third antennal segment not twice as long
as 2nd and distinctly shorter than 4th, legs short and robust,
metaibiae curved, strongly so in male. North Africa, Middle East,
Iberian Peninsula
</td>
<td class="cell_3" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20damascenum" target="_blank">
damascenum
</a>
</span>
</span>
</i>
(Fairmaire)
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
–
</td>
<td class="cell_2" colspan="1">
Third antennal segment at least twice as long as second and similar to 4th, metatibiae straight
</td>
<td class="cell_3" colspan="1">
4
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
4
</td>
<td class="cell_2" colspan="1">
Third antennal segment three times longer than 2nd. Length 7 mm. Algeria, Morocco
</td>
<td class="cell_3" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20punicum" target="_blank">
punicum
</a>
</span>
</span>
</i>
(K. & J. Daniel)
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
–
</td>
<td class="cell_2" colspan="1">
Third antennal segment at most twice longer than 2nd. Length 2.8–6mm.
</td>
<td class="cell_3" colspan="1">
5
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
5
</td>
<td class="cell_2" colspan="1">
Eyes convex, as long as temporae, pronotum as long as wide. Length 5–5.5mm. Morocco, Tunisia
</td>
<td class="cell_3" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20vaucheri" target="_blank">
vaucheri
</a>
</span>
</span>
</i>
(Vauloger)
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
–
</td>
<td class="cell_2" colspan="1">
Eyes flattened
</td>
<td class="cell_3" colspan="1">
6
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
6
</td>
<td class="cell_2" colspan="1">
Head darker than pronotum and elytra
</td>
<td class="cell_3" colspan="1">
7
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
–
</td>
<td class="cell_2" colspan="1">
Head concolorous with pronotum and elytra, body entirely yellowish brown
</td>
<td class="cell_3" colspan="1">
8
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
7
</td>
<td class="cell_2" colspan="1">
Eyes well developed, distance between
hind margin of head and hind margin of eyes at most 2 times longer than
diameter of eyes. Apical part of aedeagus short and robust, with
slightly curved ventral margin. Length 4.5–6mm. Central and southern
Europe, Turkmenistan
</td>
<td class="cell_3" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20chevrolati" target="_blank">
chevrolati
</a>
</span>
</span>
</i>
(Castelnau)
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
–
</td>
<td class="cell_2" colspan="1">
Eyes reduced, distance between hind
margin of head and hind margin of eyes at least 2.5 times longer than
diameter of eyes. Apical part of aedeagus long and narrow. Morocco
</td>
<td class="cell_3" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20angusticullum" target="_blank">
angusticullum
</a>
</span>
</span>
</i>
Hürka
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
8
</td>
<td class="cell_2" colspan="1">
Apical part of aedeagus straight, long and narrow. Length 4–5mm. Spain
</td>
<td class="cell_3" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20ramirezi" target="_blank">
ramirezi
</a>
</span>
</span>
</i>
J. and E.Vives
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
–
</td>
<td class="cell_2" colspan="1">
Apical part of aedeagus sinuate, curved, robust and hooked. Length 3.8–5.4mm. North Africa, Italy, Spain
</td>
<td class="cell_3" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20baeticum" target="_blank">
baeticum
</a>
</span>
</span>
</i>
(K. & J. Daniel)
</td>
</tr>
</tbody>
</table>
</div>
<table cellpadding="0" cellspacing="0" class="innerTable Table">
<tbody>
<tr>
<td class="cell_1" colspan="1">
1
</td>
<td class="cell_2" colspan="1">
Eyeless
</td>
<td class="cell_3" colspan="1">
2
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
–
</td>
<td class="cell_2" colspan="1">
With eyes
</td>
<td class="cell_3" colspan="1">
3
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
2
</td>
<td class="cell_2" colspan="1">
Third antennal segment only slightly
longer than 2nd and slightly shorter than 4th, anterior margin of
pronotum trapezoidal, presence of a tooth on metafemora. Length 2.7mm.
North Morocco
</td>
<td class="cell_3" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" onmouseout="hideBaloonEvent('Parazuphium_aguilerai', event)" onmouseover="showBaloon(2, 'Parazuphium_aguilerai', event)" target="_blank">
aguilerai
</a>
</span>
</span>
</i>
sp. n.
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
–
</td>
<td class="cell_2" colspan="1">
Third antennal segment more than twice
longer than 2nd and similar to 4th. Anterior margin of pronotum
bisinuate, without tooth on metafemora. Length 4.9–5.1mm. Canary
Islands
</td>
<td class="cell_3" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20feloi" onmouseout="hideBaloonEvent('Parazuphium_feloi', event)" onmouseover="showBaloon(2, 'Parazuphium_feloi', event)" target="_blank">
feloi
</a>
</span>
</span>
</i>
Machado
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
3
</td>
<td class="cell_2" colspan="1">
Third antennal segment not twice as long
as 2nd and distinctly shorter than 4th, legs short and robust,
metaibiae curved, strongly so in male. North Africa, Middle East,
Iberian Peninsula
</td>
<td class="cell_3" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20damascenum" onmouseout="hideBaloonEvent('Parazuphium_damascenum', event)" onmouseover="showBaloon(2, 'Parazuphium_damascenum', event)" target="_blank">
damascenum
</a>
</span>
</span>
</i>
(Fairmaire)
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
–
</td>
<td class="cell_2" colspan="1">
Third antennal segment at least twice as long as second and similar to 4th, metatibiae straight
</td>
<td class="cell_3" colspan="1">
4
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
4
</td>
<td class="cell_2" colspan="1">
Third antennal segment three times longer than 2nd. Length 7 mm. Algeria, Morocco
</td>
<td class="cell_3" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20punicum" onmouseout="hideBaloonEvent('Parazuphium_punicum', event)" onmouseover="showBaloon(2, 'Parazuphium_punicum', event)" target="_blank">
punicum
</a>
</span>
</span>
</i>
(K. & J. Daniel)
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
–
</td>
<td class="cell_2" colspan="1">
Third antennal segment at most twice longer than 2nd. Length 2.8–6mm.
</td>
<td class="cell_3" colspan="1">
5
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
5
</td>
<td class="cell_2" colspan="1">
Eyes convex, as long as temporae, pronotum as long as wide. Length 5–5.5mm. Morocco, Tunisia
</td>
<td class="cell_3" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20vaucheri" onmouseout="hideBaloonEvent('Parazuphium_vaucheri', event)" onmouseover="showBaloon(2, 'Parazuphium_vaucheri', event)" target="_blank">
vaucheri
</a>
</span>
</span>
</i>
(Vauloger)
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
–
</td>
<td class="cell_2" colspan="1">
Eyes flattened
</td>
<td class="cell_3" colspan="1">
6
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
6
</td>
<td class="cell_2" colspan="1">
Head darker than pronotum and elytra
</td>
<td class="cell_3" colspan="1">
7
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
–
</td>
<td class="cell_2" colspan="1">
Head concolorous with pronotum and elytra, body entirely yellowish brown
</td>
<td class="cell_3" colspan="1">
8
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
7
</td>
<td class="cell_2" colspan="1">
Eyes well developed, distance between
hind margin of head and hind margin of eyes at most 2 times longer than
diameter of eyes. Apical part of aedeagus short and robust, with
slightly curved ventral margin. Length 4.5–6mm. Central and southern
Europe, Turkmenistan
</td>
<td class="cell_3" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20chevrolati" onmouseout="hideBaloonEvent('Parazuphium_chevrolati', event)" onmouseover="showBaloon(2, 'Parazuphium_chevrolati', event)" target="_blank">
chevrolati
</a>
</span>
</span>
</i>
(Castelnau)
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
–
</td>
<td class="cell_2" colspan="1">
Eyes reduced, distance between hind
margin of head and hind margin of eyes at least 2.5 times longer than
diameter of eyes. Apical part of aedeagus long and narrow. Morocco
</td>
<td class="cell_3" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20angusticullum" onmouseout="hideBaloonEvent('Parazuphium_angusticullum', event)" onmouseover="showBaloon(2, 'Parazuphium_angusticullum', event)" target="_blank">
angusticullum
</a>
</span>
</span>
</i>
Hürka
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
8
</td>
<td class="cell_2" colspan="1">
Apical part of aedeagus straight, long and narrow. Length 4–5mm. Spain
</td>
<td class="cell_3" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20ramirezi" onmouseout="hideBaloonEvent('Parazuphium_ramirezi', event)" onmouseover="showBaloon(2, 'Parazuphium_ramirezi', event)" target="_blank">
ramirezi
</a>
</span>
</span>
</i>
J. and E.Vives
</td>
</tr>
<tr>
<td class="cell_1" colspan="1">
–
</td>
<td class="cell_2" colspan="1">
Apical part of aedeagus sinuate, curved, robust and hooked. Length 3.8–5.4mm. North Africa, Italy, Spain
</td>
<td class="cell_3" colspan="1">
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20baeticum" onmouseout="hideBaloonEvent('Parazuphium_baeticum', event)" onmouseover="showBaloon(2, 'Parazuphium_baeticum', event)" target="_blank">
baeticum
</a>
</span>
</span>
</i>
(K. & J. Daniel)
</td>
</tr>
</tbody>
</table>
</div>
<a name="phylogeneticanalysisofmoleculardata">
</a>
<span class="node_title">
Phylogenetic analysis of molecular data
</span>
<p>
The
<i>
cox1
</i>
gene fragment was aligned with no gaps,
and its correct translation to amino acids confirmed. Alignment of the
three ribosomal markers resulted in several gaps, which were included
in the analyses as obtained from MAFFT without further modifications.
Bayesian analysis reached a convergence value of 0.0005 after 20 million
generations. The initial 10% saved trees were removed as a burning
value and the half consensus tree was built with the “sumt" option in
MrBayes v.3.1.
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F5')" onmouseover="showBaloon(3, 'F5', event)">
Figure 5
</span>
represents the obtained phylogeny, were most of nodes showed very high
Bayesian posterior probabilities, which are interpreted as Bayesian
support.
</p>
<div class="imgFloatLeft">
<div class="graphicHolder">
<div class="graphicImg">
<a href="export.php_files/ZooKeys-103-049-g005.jpg" target="_blank">
<img alt="" src="export.php_files/ZooKeys-103-049-g005.jpg"/>
</a>
</div>
<div class="graphicCaptions">
<span class="graphicCaption">
</span>
<span class="graphicDescription">
</span>
</div>
<div class="unfloat">
</div>
</div>
</div>
<div class="ImgTextFloatRight">
<span class="node_label">
Figure 5.
</span>
<p>
Phylogenetic tree obtained with MrBayes for the combined dataset (
<i>
cox1, rrnL, LSU, SSU
</i>
). Numbers in nodes correspond to Bayesian posterior probabilities.
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Zuphiini" onmouseout="hideBaloonEvent('Zuphiini', event)" onmouseover="showBaloon(2, 'Zuphiini', event)" target="_blank">
Zuphiini
</a>
</span>
</span>
and
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Dryptinae" onmouseout="hideBaloonEvent('Dryptinae', event)" onmouseover="showBaloon(2, 'Dryptinae', event)" target="_blank">
Dryptinae
</a>
</span>
</span>
(sensu
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B19')" onmouseover="showBaloon(1, 'B19', event)">
Serrano 2003
</span>
) indicated with bars.
</p>
</div>
<div class="unfloat">
</div>
<div class="figureHolder" id="fig_F5">
<div class="imgFloatLeft">
<div class="graphicHolder">
<div class="graphicImg">
<a href="export.php_files/ZooKeys-103-049-g005.jpg" target="_blank">
<img alt="" src="export.php_files/ZooKeys-103-049-g005.jpg"/>
</a>
</div>
<div class="graphicCaptions">
<span class="graphicCaption">
</span>
<span class="graphicDescription">
</span>
</div>
<div class="unfloat">
</div>
</div>
</div>
<div class="ImgTextFloatRight">
<span class="node_label">
Figure 5.
</span>
<p>
Phylogenetic tree obtained with MrBayes for the combined dataset (
<i>
cox1, rrnL, LSU, SSU
</i>
). Numbers in nodes correspond to Bayesian posterior probabilities.
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Zuphiini" target="_blank">
Zuphiini
</a>
</span>
</span>
and
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Dryptinae" target="_blank">
Dryptinae
</a>
</span>
</span>
(sensu
<span class="xrefLink node_citation">
Serrano 2003
</span>
) indicated with bars.
</p>
</div>
<div class="unfloat">
</div>
</div>
<p>
We recovered a monophyletic
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Zuphiini" onmouseout="hideBaloonEvent('Zuphiini', event)" onmouseover="showBaloon(2, 'Zuphiini', event)" target="_blank">
Zuphiini
</a>
</span>
</span>
, with the two studied species of
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
</span>
</span>
</i>
as sisters, and sister to
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Zuphium" onmouseout="hideBaloonEvent('Zuphium', event)" onmouseover="showBaloon(2, 'Zuphium', event)" target="_blank">
Zuphium
</a>
</span>
</span>
</i>
(
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F5')" onmouseover="showBaloon(3, 'F5', event)">
Fig. 5
</span>
).
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Zuphiini" onmouseout="hideBaloonEvent('Zuphiini', event)" onmouseover="showBaloon(2, 'Zuphiini', event)" target="_blank">
Zuphiini
</a>
</span>
</span>
was sister to
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Drypta" onmouseout="hideBaloonEvent('Drypta', event)" onmouseover="showBaloon(2, 'Drypta', event)" target="_blank">
Drypta
</a>
</span>
</span>
</i>
, in a monophyletic
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Dryptinae" onmouseout="hideBaloonEvent('Dryptinae', event)" onmouseover="showBaloon(2, 'Dryptinae', event)" target="_blank">
Dryptinae
</a>
</span>
</span>
(sensu
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B19')" onmouseover="showBaloon(1, 'B19', event)">
Serrano 2003
</span>
).
</p>
<a name="discussion">
</a>
<span class="node_title" style="padding-top:20px; padding-bottom:0px">
Discussion
</span>
<a name="relationshipsofparazuphium">
</a>
<span class="node_title1">
Relationships of Parazuphium
</span>
<p>
The genus
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
</span>
</span>
</i>
is currently included in subtribe
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Zuphiina" onmouseout="hideBaloonEvent('Zuphiina', event)" onmouseover="showBaloon(2, 'Zuphiina', event)" target="_blank">
Zuphiina
</a>
</span>
</span>
(tribe
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Zuphiini" onmouseout="hideBaloonEvent('Zuphiini', event)" onmouseover="showBaloon(2, 'Zuphiini', event)" target="_blank">
Zuphiini
</a>
</span>
</span>
), together with
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Ildobates" onmouseout="hideBaloonEvent('Ildobates', event)" onmouseover="showBaloon(2, 'Ildobates', event)" target="_blank">
Ildobates
</a>
</span>
</span>
</i>
,
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Zuphium" onmouseout="hideBaloonEvent('Zuphium', event)" onmouseover="showBaloon(2, 'Zuphium', event)" target="_blank">
Zuphium
</a>
</span>
</span>
</i>
and
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Polistichus" onmouseout="hideBaloonEvent('Polistichus', event)" onmouseover="showBaloon(2, 'Polistichus', event)" target="_blank">
Polistichus
</a>
</span>
</span>
</i>
among the Palaearctic fauna (Baehr 2003). Although the scarcity of data
does not allow a comprehensive study, our molecular results support
this taxonomic position, with both studied
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
</span>
</span>
</i>
species clustered together as a sister group of
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Zuphium" onmouseout="hideBaloonEvent('Zuphium', event)" onmouseover="showBaloon(2, 'Zuphium', event)" target="_blank">
Zuphium
</a>
</span>
</span>
</i>
(
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F5')" onmouseover="showBaloon(3, 'F5', event)">
Fig. 5
</span>
).
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Zuphium" onmouseout="hideBaloonEvent('Zuphium', event)" onmouseover="showBaloon(2, 'Zuphium', event)" target="_blank">
Zuphium
</a>
</span>
</span>
</i>
and
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
</span>
</span>
</i>
species are recovered as related to
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Ildobates" onmouseout="hideBaloonEvent('Ildobates', event)" onmouseover="showBaloon(2, 'Ildobates', event)" target="_blank">
Ildobates
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Ildobates%20neboti" onmouseout="hideBaloonEvent('Ildobates_neboti', event)" onmouseover="showBaloon(2, 'Ildobates_neboti', event)" target="_blank">
neboti
</a>
</span>
</span>
</i>
, which was found as belonging to the
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Zuphiini" onmouseout="hideBaloonEvent('Zuphiini', event)" onmouseover="showBaloon(2, 'Zuphiini', event)" target="_blank">
Zuphiini
</a>
</span>
</span>
by
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B17')" onmouseover="showBaloon(1, 'B17', event)">
Ribera et al. (2006)
</span>
. Our data confirm the close relationship of
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Zuphium" onmouseout="hideBaloonEvent('Zuphium', event)" onmouseover="showBaloon(2, 'Zuphium', event)" target="_blank">
Zuphium
</a>
</span>
</span>
</i>
and
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
</span>
</span>
</i>
, while a more detailed phylogeny would be needed to establish the position of
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Ildobates" onmouseout="hideBaloonEvent('Ildobates', event)" onmouseover="showBaloon(2, 'Ildobates', event)" target="_blank">
Ildobates
</a>
</span>
</span>
</i>
within
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Zuphiini" onmouseout="hideBaloonEvent('Zuphiini', event)" onmouseover="showBaloon(2, 'Zuphiini', event)" target="_blank">
Zuphiini
</a>
</span>
</span>
.
</p>
<a name="classificationofparazuphium">
</a>
<span class="node_title1">
Classification of Parazuphium
</span>
<p>
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
</span>
</span>
</i>
has traditionally been divided in three subgenera,
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Neozuphium" onmouseout="hideBaloonEvent('Neozuphium', event)" onmouseover="showBaloon(2, 'Neozuphium', event)" target="_blank">
Neozuphium
</a>
</span>
</span>
</i>
, with only one valid species,
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20damascenum" onmouseout="hideBaloonEvent('Parazuphium_damascenum', event)" onmouseover="showBaloon(2, 'Parazuphium_damascenum', event)" target="_blank">
damascenum
</a>
</span>
</span>
</i>
(
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B12')" onmouseover="showBaloon(1, 'B12', event)">
Mateu 1988
</span>
) (note that Baehr 2003 does not include the synonymisation of
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20varum" onmouseout="hideBaloonEvent('Parazuphium_varum', event)" onmouseover="showBaloon(2, 'Parazuphium_varum', event)" target="_blank">
varum
</a>
</span>
</span>
</i>
,
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20bactrianum" onmouseout="hideBaloonEvent('Parazuphium_bactrianum', event)" onmouseover="showBaloon(2, 'Parazuphium_bactrianum', event)" target="_blank">
bactrianum
</a>
</span>
</span>
</i>
and
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20novaki" onmouseout="hideBaloonEvent('Parazuphium_novaki', event)" onmouseover="showBaloon(2, 'Parazuphium_novaki', event)" target="_blank">
novaki
</a>
</span>
</span>
</i>
by
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B12')" onmouseover="showBaloon(1, 'B12', event)">
Mateu 1988
</span>
);
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
</span>
</span>
</i>
s.str., with 12 Palaearctic (Baehr 2003) and 20 Ethiopian species; and
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Austrozuphium" onmouseout="hideBaloonEvent('Austrozuphium', event)" onmouseover="showBaloon(2, 'Austrozuphium', event)" target="_blank">
Austrozuphium
</a>
</span>
</span>
</i>
, with five Australasian species (
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B2')" onmouseover="showBaloon(1, 'B2', event)">
Baehr 1985
</span>
,
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B9')" onmouseover="showBaloon(1, 'B9', event)">
Lorenz 2005
</span>
). The latter species are of dubious affiliation, and maybe not directly related to the Palaearctic and Ethiopian species (
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B13')" onmouseover="showBaloon(1, 'B13', event)">
Mateu 1993
</span>
and pers. comm. 2008).
</p>
<p>
The subgenus
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Neozuphium" onmouseout="hideBaloonEvent('Neozuphium', event)" onmouseover="showBaloon(2, 'Neozuphium', event)" target="_blank">
Neozuphium
</a>
</span>
</span>
</i>
was described by
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B5')" onmouseover="showBaloon(1, 'B5', event)">
Hürka (1982)
</span>
based on the relative length of the 2nd to 4th antennomeres and the
shape of the legs, more robust and with curved tibia and enlarged
femora in the males in
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Neozuphium" onmouseout="hideBaloonEvent('Neozuphium', event)" onmouseover="showBaloon(2, 'Neozuphium', event)" target="_blank">
Neozuphium
</a>
</span>
</span>
</i>
.
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" onmouseout="hideBaloonEvent('Parazuphium_aguilerai', event)" onmouseover="showBaloon(2, 'Parazuphium_aguilerai', event)" target="_blank">
aguilerai
</a>
</span>
</span>
</i>
sp. n. has the third antennomere only slightly longer than the 2nd and slightly shorter than the 4th (
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F2')" onmouseover="showBaloon(3, 'F2', event)">
Fig. 2e
</span>
), so it would agree with
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Neozuphium" onmouseout="hideBaloonEvent('Neozuphium', event)" onmouseover="showBaloon(2, 'Neozuphium', event)" target="_blank">
Neozuphium
</a>
</span>
</span>
</i>
(species of
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
</span>
</span>
</i>
s.str. have the 3rd antennomere double than the 2nd, and similar to the 4th,
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B5')" onmouseover="showBaloon(1, 'B5', event)">
Hürka 1982
</span>
,
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B12')" onmouseover="showBaloon(1, 'B12', event)">
Mateu 1988
</span>
). However, the shape and size of the legs do not agree with the diagnostic characters of
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Neozuphium" onmouseout="hideBaloonEvent('Neozuphium', event)" onmouseover="showBaloon(2, 'Neozuphium', event)" target="_blank">
Neozuphium
</a>
</span>
</span>
</i>
, as the males have straight metatibia (
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F2')" onmouseover="showBaloon(3, 'F2', event)">
Fig. 2m
</span>
) and they are in general slender and long in comparison to
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Neozuphium" onmouseout="hideBaloonEvent('Neozuphium', event)" onmouseover="showBaloon(2, 'Neozuphium', event)" target="_blank">
(Neozuphium)
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20%28Neozuphium%29%20damascenum" onmouseout="hideBaloonEvent('Parazuphium_(Neozuphium)_damascenum', event)" onmouseover="showBaloon(2, 'Parazuphium_(Neozuphium)_damascenum', event)" target="_blank">
damascenum
</a>
</span>
</span>
</i>
(
<span class="xrefLink node_fig" onmouseout="hideBaloonEvent('F2')" onmouseover="showBaloon(3, 'F2', event)">
Figs 2i-n
</span>
).
These are, in any case, characters with dubious phylogenetic
information, so instead of redefining the subgenera, or describing
additional taxa, we opt to follow
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B19')" onmouseover="showBaloon(1, 'B19', event)">
Serrano (2003)
</span>
, treating the subgenus
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Neozuphium" onmouseout="hideBaloonEvent('Neozuphium', event)" onmouseover="showBaloon(2, 'Neozuphium', event)" target="_blank">
Neozuphium
</a>
</span>
</span>
</i>
Hürka as consubgeneric with
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
</span>
</span>
</i>
Jeannel, and the former name as a junior synonym of the latter.
</p>
<a name="endogeanwayoflifeinzuphiini">
</a>
<span class="node_title1">
Endogean way of life in Zuphiini
</span>
<p>
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" onmouseout="hideBaloonEvent('Parazuphium_aguilerai', event)" onmouseover="showBaloon(2, 'Parazuphium_aguilerai', event)" target="_blank">
aguilerai
</a>
</span>
</span>
</i>
sp. n. differs from all other known species of the genus in its clear
adaptations to an endogean way of life. Other species are regularly
found in soil crevices, specially among the cracks of the dried
substratum of areas which are regularly inundated (
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B2')" onmouseover="showBaloon(1, 'B2', event)">
Baehr 1985
</span>
,
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B10')" onmouseover="showBaloon(1, 'B10', event)">
Machado 1992
</span>
,
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B7')" onmouseover="showBaloon(1, 'B7', event)">
Lencina and Serrano 1995
</span>
).
These species have some modifications suggesting an adaptation to this
cryptic way of life (small size, flattened body, some degree of
depigmentation, microphthalmy,
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B6')" onmouseover="showBaloon(1, 'B6', event)">
Jeannel 1942
</span>
), but not to the extent of
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" onmouseout="hideBaloonEvent('Parazuphium_aguilerai', event)" onmouseover="showBaloon(2, 'Parazuphium_aguilerai', event)" target="_blank">
aguilerai
</a>
</span>
</span>
</i>
sp. n., which was found in company of other typical endogean insects (
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Leptanilla" onmouseout="hideBaloonEvent('Leptanilla', event)" onmouseover="showBaloon(2, 'Leptanilla', event)" target="_blank">
Leptanilla
</a>
</span>
</span>
</i>
sp.,
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Amblyopone" onmouseout="hideBaloonEvent('Amblyopone', event)" onmouseover="showBaloon(2, 'Amblyopone', event)" target="_blank">
Amblyopone
</a>
</span>
</span>
</i>
sp.,
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Torneuma" onmouseout="hideBaloonEvent('Torneuma', event)" onmouseover="showBaloon(2, 'Torneuma', event)" target="_blank">
Torneuma
</a>
</span>
</span>
</i>
sp.) below a deeply buried large stone in a hillside. The only other anophthalmous species of the genus (
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20feloi" onmouseout="hideBaloonEvent('Parazuphium_feloi', event)" onmouseover="showBaloon(2, 'Parazuphium_feloi', event)" target="_blank">
feloi
</a>
</span>
</span>
</i>
) was found in a cave, and it is larger and with longer appendages (
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B11')" onmouseover="showBaloon(1, 'B11', event)">
Machado 1998
</span>
), as is typical of cave fauna inhabiting larger open spaces.
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" onmouseout="hideBaloonEvent('Parazuphium_aguilerai', event)" onmouseover="showBaloon(2, 'Parazuphium_aguilerai', event)" target="_blank">
aguilerai
</a>
</span>
</span>
</i>
sp. n. shares with
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20feloi" onmouseout="hideBaloonEvent('Parazuphium_feloi', event)" onmouseover="showBaloon(2, 'Parazuphium_feloi', event)" target="_blank">
feloi
</a>
</span>
</span>
</i>
,
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20chevrolati" onmouseout="hideBaloonEvent('Parazuphium_chevrolati', event)" onmouseover="showBaloon(2, 'Parazuphium_chevrolati', event)" target="_blank">
chevrolati
</a>
</span>
</span>
</i>
and
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20vaucheri" onmouseout="hideBaloonEvent('Parazuphium_vaucheri', event)" onmouseover="showBaloon(2, 'Parazuphium_vaucheri', event)" target="_blank">
vaucheri
</a>
</span>
</span>
</i>
the presence of a spine in the metatibia (
<span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B11')" onmouseover="showBaloon(1, 'B11', event)">
Machado 1998
</span>
), although at the moment it is not possible to assert the phylogenetic value of this character.
</p>
</div>
<a name="acknowledgements">
</a>
<div class="smallSecHolder acknowledgementsHolder">
<div class="acknowledgementsText">
<span class="node_title">
Acknowledgements
</span>
<p>
This research was supported in part by the project
CGL2007-61665 of the Spanish Ministry of Science and Innovation and the
project S/REF.08724/PI/08 of the Fundacion Seneca. CA had a FPU
predoctoral grant of the same Ministry. Thanks are due to José Serrano
and José Luis Lencina for specimens of
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20damascenum" onmouseout="hideBaloonEvent('Parazuphium_damascenum', event)" onmouseover="showBaloon(2, 'Parazuphium_damascenum', event)" target="_blank">
damascenum
</a>
</span>
</span>
</i>
and
<i>
<span class="node_taxon-name">
<span>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">
Parazuphium
</a>
<a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20ramirezi" onmouseout="hideBaloonEvent('Parazuphium_ramirezi', event)" onmouseover="showBaloon(2, 'Parazuphium_ramirezi', event)" target="_blank">
ramirezi
</a>
</span>
</span>
</i>
used in comparisons and for the help in the identification of some
species. Joaquin Mateu, Arnaud Faille and two anonymous Referees helped
with their comments. Jesús Arribas did the drawing of habitus.
Phylogenetic analyses were carried out on the freely available Bioportal
(www.bioportal.uio.no)
</p>
</div>
</div>
<a name="references">
</a>
<div class="smallSecHolder referencesHolder">
<div class="referencesText">
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Line drawing ofhabitus of Parazuphium aguilerai sp. n. Total length 2.7 mm.
In [7]:
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print(figcap)
[<div class="ImgTextFloatRight">
<span class="node_label">Figure 1.</span>
<p>Line drawing ofhabitus of <i><span class="node_taxon-name"><span><a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">Parazuphium</a> <a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" onmouseout="hideBaloonEvent('Parazuphium_aguilerai', event)" onmouseover="showBaloon(2, 'Parazuphium_aguilerai', event)" target="_blank">aguilerai</a></span></span></i> sp. n. Total length 2.7 mm.</p>
</div>, <div class="ImgTextFloatRight">
<span class="node_label">Figure 1.</span>
<p>Line drawing ofhabitus of <i><span class="node_taxon-name"><span><a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" target="_blank">Parazuphium</a> <a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" target="_blank">aguilerai</a></span></span></i> sp. n. Total length 2.7 mm.</p>
</div>, <div class="ImgTextFloatRight">
<span class="node_label">Figure 2.</span>
<p>Photographic images of<i><span class="node_taxon-name"><span><a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">Parazuphium</a> <a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" onmouseout="hideBaloonEvent('Parazuphium_aguilerai', event)" onmouseover="showBaloon(2, 'Parazuphium_aguilerai', event)" target="_blank">aguilerai</a></span></span></i> sp. n. <b>A</b> whole specimen <b>B</b> head in dorso-lateral view <b>C</b> labial palpus; (d), maxillary palpus <b>C</b> antenna <b>F</b> margin of left elytron in lateral view <b>G</b> margin of right elytron, detail for anterior umbilicate setae, numbers 1 to 5 <b>H</b> margin of right elytron, detail of posterior umbilicate setae, numbers 6 to10, arrows over them point other smaller setae <b>I–N</b> details of anterior, median and posterior legs respectively.</p>
</div>, <div class="ImgTextFloatRight">
<span class="node_label">Figure 2.</span>
<p>Photographic images of<i><span class="node_taxon-name"><span><a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" target="_blank">Parazuphium</a> <a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" target="_blank">aguilerai</a></span></span></i> sp. n. <b>A</b> whole specimen <b>B</b> head in dorso-lateral view <b>C</b> labial palpus; (d), maxillary palpus <b>C</b> antenna <b>F</b> margin of left elytron in lateral view <b>G</b> margin of right elytron, detail for anterior umbilicate setae, numbers 1 to 5 <b>H</b> margin of right elytron, detail of posterior umbilicate setae, numbers 6 to10, arrows over them point other smaller setae <b>I–N</b> details of anterior, median and posterior legs respectively.</p>
</div>, <div class="ImgTextFloatRight">
<span class="node_label">Figure 3.</span>
<p>Photographic image of median lobe of <i><span class="node_taxon-name"><span><a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">Parazuphium</a> <a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" onmouseout="hideBaloonEvent('Parazuphium_aguilerai', event)" onmouseover="showBaloon(2, 'Parazuphium_aguilerai', event)" target="_blank">aguilerai</a></span></span></i> sp. n. Left lateral aspect.</p>
</div>, <div class="ImgTextFloatRight">
<span class="node_label">Figure 3.</span>
<p>Photographic image of median lobe of <i><span class="node_taxon-name"><span><a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" target="_blank">Parazuphium</a> <a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" target="_blank">aguilerai</a></span></span></i> sp. n. Left lateral aspect.</p>
</div>, <div class="ImgTextFloatRight">
<span class="node_label">Figure 4.</span>
<p>Habitat of <i><span class="node_taxon-name"><span><a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" onmouseout="hideBaloonEvent('Parazuphium', event)" onmouseover="showBaloon(2, 'Parazuphium', event)" target="_blank">Parazuphium</a> <a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" onmouseout="hideBaloonEvent('Parazuphium_aguilerai', event)" onmouseover="showBaloon(2, 'Parazuphium_aguilerai', event)" target="_blank">aguilerai</a></span></span></i> sp. n.</p>
</div>, <div class="ImgTextFloatRight">
<span class="node_label">Figure 4.</span>
<p>Habitat of <i><span class="node_taxon-name"><span><a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium" target="_blank">Parazuphium</a> <a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Parazuphium%20aguilerai" target="_blank">aguilerai</a></span></span></i> sp. n.</p>
</div>, <div class="ImgTextFloatRight">
<span class="node_label">Figure 5.</span>
<p>Phylogenetic tree obtained with MrBayes for the combined dataset (<i>cox1, rrnL, LSU, SSU</i>). Numbers in nodes correspond to Bayesian posterior probabilities. <span class="node_taxon-name"><span><a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Zuphiini" onmouseout="hideBaloonEvent('Zuphiini', event)" onmouseover="showBaloon(2, 'Zuphiini', event)" target="_blank">Zuphiini</a></span></span> and <span class="node_taxon-name"><span><a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Dryptinae" onmouseout="hideBaloonEvent('Dryptinae', event)" onmouseover="showBaloon(2, 'Dryptinae', event)" target="_blank">Dryptinae</a></span></span> (sensu <span class="xrefLink node_citation" onmouseout="hideBaloonEvent('B19')" onmouseover="showBaloon(1, 'B19', event)">Serrano 2003</span>) indicated with bars.</p>
</div>, <div class="ImgTextFloatRight">
<span class="node_label">Figure 5.</span>
<p>Phylogenetic tree obtained with MrBayes for the combined dataset (<i>cox1, rrnL, LSU, SSU</i>). Numbers in nodes correspond to Bayesian posterior probabilities. <span class="node_taxon-name"><span><a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Zuphiini" target="_blank">Zuphiini</a></span></span> and <span class="node_taxon-name"><span><a class="taxonNameLink" href="http://ptp.pensoft.eu/external_details.php?type=1&query=Dryptinae" target="_blank">Dryptinae</a></span></span> (sensu <span class="xrefLink node_citation">Serrano 2003</span>) indicated with bars.</p>
</div>]
In [33]:
dirname = ("1124-G-1-layout.html")
chomped = dirname[:-5]
string = re.sub('[^0-9]', '', dirname[:-14])
print string
print dirname
print chomped
1124
1124-G-1-layout.html
1124-G-1-layout
In [152]:
file = open("./"+chomped+"/"+chomped+".tmp", "wb")
In [153]:
for tr in figcap[1::2]:
cap = (tr.get_text()).strip().encode("utf8")
#print cap
file.write(cap+'\n')
In [154]:
file.flush()
file.close()
In [157]:
figimg = soup.find_all('div', class_="graphicImg")
print(figimg)
[<div class="graphicImg"><a href="export.php_files/ZooKeys-103-049-g001.jpg" target="_blank"><img alt="" src="export.php_files/ZooKeys-103-049-g001.jpg"/></a></div>, <div class="graphicImg"><a href="export.php_files/ZooKeys-103-049-g001.jpg" target="_blank"><img alt="" src="export.php_files/ZooKeys-103-049-g001.jpg"/></a></div>, <div class="graphicImg"><a href="export.php_files/ZooKeys-103-049-g002.jpg" target="_blank"><img alt="" src="export.php_files/ZooKeys-103-049-g002.jpg"/></a></div>, <div class="graphicImg"><a href="export.php_files/ZooKeys-103-049-g002.jpg" target="_blank"><img alt="" src="export.php_files/ZooKeys-103-049-g002.jpg"/></a></div>, <div class="graphicImg"><a href="export.php_files/ZooKeys-103-049-g003.jpg" target="_blank"><img alt="" src="export.php_files/ZooKeys-103-049-g003.jpg"/></a></div>, <div class="graphicImg"><a href="export.php_files/ZooKeys-103-049-g003.jpg" target="_blank"><img alt="" src="export.php_files/ZooKeys-103-049-g003.jpg"/></a></div>, <div class="graphicImg"><a href="export.php_files/ZooKeys-103-049-g004.jpg" target="_blank"><img alt="" src="export.php_files/ZooKeys-103-049-g004.jpg"/></a></div>, <div class="graphicImg"><a href="export.php_files/ZooKeys-103-049-g004.jpg" target="_blank"><img alt="" src="export.php_files/ZooKeys-103-049-g004.jpg"/></a></div>, <div class="graphicImg"><a href="export.php_files/ZooKeys-103-049-g005.jpg" target="_blank"><img alt="" src="export.php_files/ZooKeys-103-049-g005.jpg"/></a></div>, <div class="graphicImg"><a href="export.php_files/ZooKeys-103-049-g005.jpg" target="_blank"><img alt="" src="export.php_files/ZooKeys-103-049-g005.jpg"/></a></div>]
In [162]:
file = open("./"+chomped+"/"+chomped+"-imgs.txt", "wb")
In [163]:
string = re.sub('[^0-9]', '', dirname[:-14])
for tr in figimg[1::2]:
imgs = tr.find('a')['href']
#print ("http://www.pensoft.net/J_FILES/1/articles/"+string+"/"+imgs)
file.write("http://www.pensoft.net/J_FILES/1/articles/"+string+"/"+imgs+'\n')
In [164]:
file.flush()
file.close()
In [187]:
ayt = soup.find_all('div', class_="authors")
print (ayt)
---------------------------------------------------------------------------
AttributeError Traceback (most recent call last)
<ipython-input-187-9a1e24c2f712> in <module>()
1 ayt = soup.find_all('div', class_="authors")
----> 2 shit = ayt.encode("utf8")
3 print (shit)
AttributeError: 'ResultSet' object has no attribute 'encode'
In [145]:
file = open("./"+chomped+"/bibinfo.txt", "wb")
In [196]:
from string import digits
tit = soup.find('title')
title = tit.string.strip().encode("utf8")
doibit = soup.find('div', class_="doiNumber")
outit = (doibit.string).strip().replace("doi: ","http://dx.doi.org/").encode("utf8")
blah = unicode(outit.strip().replace("doi: ","http://dx.doi.org/"),'utf-8')
#print (title+". "+outit)
#file.write(title+". "+outit)
for bl in ayt:
authors = (bl.get_text()).encode("utf8")
cheese = (authors.translate(None, digits)).replace("†","").replace("‡","").replace("§","").replace("|","").replace("¶","").replace("#","").replace(",,",",")
#print (re.sub('[\W_]+', ' ', authors)) #Carmelo And jar1 Carles Hernando2 Ignacio Ribera3
print (unicode(cheese,'utf-8')+" "+title+". "+blah)
#file.write("Source: "+cheese+" "+title+". "+outit)
Carmelo Andújar, Carles Hernando, Ignacio Ribera,
A new endogean, anophthalmous species of Parazuphium Jeannel from Northern Morocco (Coleoptera, Carabidae), with new molecular data for the tribe Zuphiini. ZooKeys 103: 49–62, http://dx.doi.org/10.3897/ZooKeys.103.1124
In [147]:
file.flush()
file.close()
In [168]:
doibit = soup.find('div', class_="doiNumber")
print (doibit.string)
ZooKeys 103: 49–62, doi: 10.3897/ZooKeys.103.1124
In [133]:
file = open("./"+chomped+"/bibinfo.txt", "wb")
In [139]:
tit = soup.find('title')
title = tit.string.strip().encode("utf8")
doibit = soup.find('div', class_="doiNumber")
outit = (doibit.string).strip().replace("doi: ","http://dx.doi.org/").encode("utf8")
print (title+". "+outit)
#file.write(title+". "+outit)
A new endogean, anophthalmous species of Parazuphium Jeannel from Northern Morocco (Coleoptera, Carabidae), with new molecular data for the tribe Zuphiini. ZooKeys 103: 49–62, http://dx.doi.org/10.3897/ZooKeys.103.1124
In [135]:
file.flush()
file.close()
In [326]:
file = open("./"+chomped+"/bibinfo.txt", "wb")
In [327]:
desc = soup.findAll(attrs={"name":"dc.title"})
titll = desc[0]['content'].encode('utf-8')
desc = soup.findAll(attrs={"name":"dc.source"})
bibinfo = desc[0]['content'].encode('utf-8')
#print bibinfo
#print titll
file.write(titll+" "+bibinfo)
In [328]:
file.flush()
file.close()
In [173]:
tit = soup.find('title')
title = tit.string.strip().encode("utf8")
print title
A new endogean, anophthalmous species of Parazuphium Jeannel from Northern Morocco (Coleoptera, Carabidae), with new molecular data for the tribe Zuphiini
In [180]:
chep = soup.title(text=True)
print (chep.string)
---------------------------------------------------------------------------
AttributeError Traceback (most recent call last)
<ipython-input-180-b4190a0cca64> in <module>()
1 chep = soup.title(text=True)
----> 2 print (chep.string)
AttributeError: 'ResultSet' object has no attribute 'string'
In [186]:
desc = soup.findAll("title")
first = desc[0]
proc = first.replace("<title>","").replace("</title>","")
---------------------------------------------------------------------------
TypeError Traceback (most recent call last)
<ipython-input-186-3f853caae301> in <module>()
1 desc = soup.findAll("title")
2 first = desc[0]
----> 3 proc = first.replace("<title>","").replace("</title>","")
TypeError: 'NoneType' object is not callable
In [ ]:
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